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Sheep (Ovis aries) resting on the Quiraing mountains with low cloud cover, Isle of Skye, Inner Hebrides, Scotland, UK. July
Close up of European dark bee (Apis mellifera mellifera - nigra race), feeding on ice plant flowers (Sedum spectabile), Monmouthshire, Wales, UK. September
Transparent ice formation on Lake Baikal, Siberia, Russia, March 2012
Red-crowned Crane (Grus japonensis) preening in snow, Hokkaido, Japan, February
Grasshopper Warbler (Locustella naevia) singing, Joensuu, Finland, May
Capercaillie (Tetrao urogallus) male in flight, Jalasjarvi, Finland, April
Young rabbit hiding in grass, Okunoshima Rabbit island, Takehara, Hiroshima, Japan
Quaking aspen trees (Populus tremuloides) and conifers with dusting of snow, Dome Hill above Dawson City, Yukon Territories, Canada, September 2013
Green waters of Emerald Lake, near Carcross, Yukon Territories, Canada, September 2013
Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya
Spotted reed frog (Hyperolius pucticulatus) tadpoles, captive, endemic to Tanzania
European hornet (Vespa crabro) worker in flight, Surrey, England, August
The Pont St-Benezet, Palais des Papes and Rhone River at dusk, Avignon, Provence, France, October 2012
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Sea otter (Enhydra lutris) floating on its back at surface among kelp, Alaska, USA. Gulf of Alaska, Pacific ocean
Ground squirrels (Xerus inauris ) one with its tail over the back of another, Kgalagadi Transfrontier Park, Northern Cape, South Africa, February
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Pair of Blesbok (Damaliscus pygargus) galloping across the open flood plains of the Bushmans River, Kariega Game Reserve, South Africa. Overall winner of the GDT Competition 2014
Wild Mustang horses running, Sand Wash Basin Herd Area, Colorado, USA
Wild Mustang horses fighting McCullough Peaks Herd Area, Wyoming, USA
Fallow deer (Dama dama) buck crossing road under street lights. London, UK. January
Greylag geese (Anser anser). Cley, Norfolk, UK, March. Digital composite
Silvery-cheeked Hornbill (Bycanistes brevis) close-up at zoo, occurs in East Africa
African fish eagle (Haliaeetus vocifer) in flight, Chobe National Park, Botswana
Samoyed dog running in in snow, Ledyard, Connecticut, USA
Sperm whale (Physeter macrocephalus) tail fluke above water during dive, Kaikoura, New Zealand, July, Vulnerable species
Four Cedar waxwings (Bombycilla cedrorum) feeding on Crabapple (Malus sp) fruit, Ithaca, New York, USA, February
Brazilian tanager (Ramphocelus bresilius) perched on branch, Serrinha do Alambari, Penedo, Itatiaia, Rio de Janeiro, Brazil
Red-tailed bumblebee (Bombus lapidarius) taking off from Cornflower (Centaurea cyanea) in Bishops Meadow, Farnham. Surrey, England, UK, July 2014. Digital composite
Male Pheasant (Phasianus colchicus) flapping wings, Hungary, June
Cowslips (Primula veris) on conservation headland on organic farm. Norfolk, UK, April
Chestnut Mustang running in snow, at ranch, Shell, Wyoming, USA. February
Red and green macaws (Ara chloropterus) in flight, motion blurred photograph, Buraxo das aras, Brazil
Adult Helmet Vanga (Euryceros prevostii) incubating a clutch of 3 eggs. Nest constructed in the fork of a Birds Nest Fern (Asplenium sp.) Masoala National Park, north east Madagascar
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in March. Wapusk National Park. Churchill, Manitoba, Canada
Masai Giraffe (Giraffa camelopardalis tippelskirchi) juveniles, Masai Mara Game Reserve, Kenya
Vancouver Island black bear (Ursus americanus vancouveri) mother with cubs on a beach, Vancouver Island, British Columbia, Canada, July
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
Adult male Diademed Sifaka (Propithecus diadema) skipping / dancing across open ground between forest fragments, eastern Madagascar
Temperate rainforest with Vine maple (Acer circinatum) and fern, Golden Ears provincial park, British Columbia, Canada, July
Great Basin Bristlecone Pine (Pinus longaeva) fallen ancient tree, White Mountains, California, USA, March
Wood Anemones (Anemone nemorosa) in flower on woodland floor, low angle fish eye view, Peak District National Park, Derbyshire, UK, May
Yacare Caiman (Caiman yacare) with butterfly (Paulogramma pyracmon) resting on its snout, Pantanal, Brazil
Female Jaguar (Panthera onca) playing with her cub, captive, occurs in Southern and Central America
Black Lemur (Eulemur macaco) male, Nosy Komba, Madagascar
Pika (Ochotona princeps) in scree rock pile, Sheepeaters Cliff, Yellowstone National Park, Wyoming, USA, September
Black-and-white kitten standing, against white background DIGITALLY ENHANCED
Harvest mouse (Micromys minutus) on wheat stem feeding, Devon, UK, July. Captive