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Carp (Cyprinus carpio) Aare River, Switzerland 2006
Killer Whales / Orcas (Orcinus orca)Killer whales / Orcas (Orcinus orca) large pod including calf traveling together while foraging on large schools of Herring (Clupea harengus) in the cold waters of northern Norway, January
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef
Leafy seadragon (Phycodurus eques) male carrying eggs, swims over seagrass meadow. Wool Bay, Edithburgh, South Australia. Gulf of St Vincent
Tompot blenny (Parablennius gattorugine) in bright summer mating colours, peering out from his home in a discarded tube, where he was guarding his eggs (not visible in this photo), Swanage Pier
Northern pike (Esox lucius) amongst Shining pondweed (Potamogeton lucens). Lake Neuchatel, Switzerland, September. Photographed for The Freshwater Project
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
Female Anglerfish fanfin seadevil (Caulophryne jordani) - Deep sea species from Atlantic Ocean, found at 700-3000m
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Weedy seadragon (Phyllopteryx taeniolatus) male carries eggs through a kelp forest (Macrocystis pyrifera) in Tasmania, Australia. Tasmania is the only part of Australia with giant kelp forests
School of Blackfin barracuda (Sphyraena qenie) forming circle in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Hairy frogfish / Striated frogfish (Antennarius striatus), portrait, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
Inflated spotted porcupinefish (Diodon hystrix), Maui, Hawaii
Brown trout (Salmo trutta) leaping for a damselfly. (digitally enhanced)
Bat fish juvenile {Halicutaea sp} deep sea
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
Atlantic halibut (Hippoglossus hippoglossus) portrait, Saltstraumen, Bod, Norway, October 2008
A portrait of a Mediterranean parrotfish (Sparisoma cretense) above an algae covered rock. El Hierro, Canary Islands, Spain. East Atlantic Ocean
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Illuminated netdevil (Linophyne arboritera), male anglerfish attached to much larger female, captive from deep sea
Portrait of Ballan wrasse (Labrus bergylta), St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, October 2011
Arctic charr (Salvelinus alpinus) males showing breeding colours, in a river where they have come to spawn, Ennerdale Valley, Lake District NP, Cumbria, England, UK, November 2011
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Yellow and blueback fusilier (Caesio teres) shoal, Similan Islands, Andaman Sea, Thailand
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Lined seahorses (Hippocampus erectus) amongst corals, The Bahamas
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Brown trout (Salmo trutta) in shallow water migrating upstream, Bornholm, Denmark, November 2008
Bream (Abramis brama) and European chub (Leuciscus / Squalius cephalus) shoal in a calm part of the Rhine, Schaffhausen, Switzerland, February 2009 WWE OUTDOOR EXHIBITION
Pike (Esox lucius) in disused quarry, Stoney Stanton, Stoney Cove, Leicestershire, UK, June
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp)Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Healthy coral reef landscape, Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Archerfish jets water at insect to dislodge it from branch {Toxotes chatareus} from SE Asia
Yellow tropical reef fish composite image on black background, Falcula butterflyfish (Chaetodon falcula), Yellow boxfish (Ostracion cubicus), Golden butterflyfish (Chaetodon semilarvatus)
Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata)
West African mudskipper (Periophthalmus barbarus) Joao Vieira Poillo, Bijagos Archipellago. Guinea Bissau
Three Sockeye salmon/ Red salmon (Oncorhynchus nerka) swimming upstream as they migrate back to the river of their birth to spawn
Tail-spot blenny (Ecsenius stigmatura) peering out from amongst the coral, Triton Bay, West Papua, Indonesia, Pacific Ocean
Guineafowl puffer fish (Arothron meleagris) inflated, displaying defensive behaviour over coral reef, Yap, Indo Pacific, Micronesia
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) shoal swimming over Turtlegrass (Halophila stipulacea) seagrass bed. Marsa Alam, Egypt
Deep sea fish (Winteria telescopa) with forward pointing tubular eyes
Short-snouted seahorse (Hippocampus hippocampus) male, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean Sea
Giant hogfish (Bodianus macrognathos) male yawning at a cleaning station run by juvenile Spanish hogfish (Bodianus rufus). Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Black grouper (Mycteroperca bonaci) yawning on a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Yellowfin tuna (Thunnus albacares) hunting mackerel (Scomber colias) Tenerife, Canary Islands
Unicorn leatherjacket filefish (Aluterus monoceros) at night, The Bahamas. February
Tiger moray / Fangtooth Moray (Enchelycore anatina) and Black moray eel (Muraena augusti), South Tenerife, Canary Islands, Atlantic Ocean
Couple of Blue shark (Prionace glauca) and Pilot fish (Naucrates ductor), Pico Island, Azores, Portugal, Atlantic Ocean
Blue shark (Prionace glauca) with fishing hook caught in mouth and Pilot fish (Naucrates ductor), Pico Island, Azores, Portugal, Atlantic Ocean
Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands
Ocean sunfish (Mola mola) and Pilot fish (Naucrates sp). Tenerife, Canary Islands
Napoleon wrasse (Cheilinus undulatus) on a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Chinese sturgeon (Acipenser sinensis), captive in aquarium. Adults are no longer able to swim upriver to spawn due to Gezhouba and Three Gorges dams
Bubble eye goldfish (Carassius auratus) with upward pointing eyes and two large fluid-filled sacs.. In captivity, Shanghai Zoo, Shanghai, China
Chinese grass carp / White amur (Ctenopharyngodon idella), farmed for food in China. Captive specimen, China
Golden grouper (Mycteroperca rosacea), yellow color-variation of the most common leopard grouper species, Cabo Pulmo Marine National Park, Baja California Sur, Mexico
Blue shark (Prionace glauca) with a pair of pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Moorish idol (Zanclus cornutus) juvenile with transparent skin in the open ocean at night off Anilao, Philippines
Pelagic thresher shark (Alopias pelagicus) being cleaned by a bluestreak cleaner wrasse (Labroides dimidiatus) at Monad Shoal off Malapascua Island, Philippines
Ribbon eel ( Rhinomuraena quaesita) Lembeh Strait, North Sulawesi, Indonesia
Taylors pygmy leatherjacket (Brachaluteres taylori). Lembeh Strait, North Sulawesi, Indonesia
An adult female Spiny Seahorse (Hippocampus guttulatus) in a meadow of seagrass (Zostera marina). Studland Bay, Dorset, England, UK, August
Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico
