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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, September
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
Killer Whales / Orcas (Orcinus orca)Killer whales / Orcas (Orcinus orca) large pod including calf traveling together while foraging on large schools of Herring (Clupea harengus) in the cold waters of northern Norway, January
Butt of Lewis lighthouse, Port Nis / Port of Ness, Isle of Lewis, Outer Hebrides, Scotland, UK. March, 2015
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Humpback whale (Megaptera novaeangliae) breaching, near Hout Bay, South Africa
Sea stacks, Duncansby Head, John O Groats, Caithness, Scotland, UK, April 2015
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, July
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Orca whales (Orcinus orca) pod surfacing together, Shetland, Scotland, UK. April
Grey seal (Halichoerus grypus) swimming towards camera, Orkney, Scotland, UK, August
Gannet (Morus bassanus) adult turns in the air as it begins to plunge dive. Shetland Islands, Scotland, UK, July
Weedy seadragon (Phyllopteryx taeniolatus) male carries eggs through a kelp forest (Macrocystis pyrifera) in Tasmania, Australia. Tasmania is the only part of Australia with giant kelp forests
Wandering albatross {Diomedea exulans} flying over open ocean, South Atlantic
Florida manatee (Trichechus manatus latirostris) mother and calf endangered. Three Sisters Spring in Crystal River, Florida, USA
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Sea otter (Enhydra Lutris) with forelegs raised, Sitka Sound, Alaska, USA, August
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
RF - Composite image of tropical nudibranchs on a black background showing variety and abundance of nudibranch species, Indo-Pacific
Deep sea jellyfish (Periphylla periphylla) juvenile, Trondheimsfjord, Norway, Atlantic ocean
View of Boscastle harbour and Cornish coastline. North Cornwall, UK, June 2010. NOT AVAILABLE FOR BOOK COVER USE UNTIL OCTOBER 2024 WORLDWIDE
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
Dolphin trainer interacting with Bottlenose Dolphin (Tursiops truncatus), Dolphin Reef, Eilat, Israel - Red Sea. Model released
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Angular roughshark (Oxynotus centrina) a deepsea species living at 80-300m depth, caught by fishermen and released, Costa Brava, Catalunya, Spain, Mediterranean Sea
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Roughtail stingray (Dasyatis centura) on sea floor, Tenerife, Canary Islands
Electric fileclam / Disco clam (Ctenoides ales). West Papua, Indonesia. Indo-West Pacific
Humpback whale (Megaptera novaeangliae) male calf relaxing with his mother and an escort whale in the background. Vava'u, Tonga, South Pacific
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
Humpback whale (Megaptera novaeangliae) just under surface, off Shetland, Scotland, UK. First ever underwater images of humpbacks in British waters