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Hooded seal male display {Cystophora cristata} St Lawrence gulf, Canada
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Proboscis monkey (Nasalis larvatus) male, Sabah, Borneo
Wild boar (Sus scrofa) mother watching over tiny piglets, the Netherlands, April
RF- Close-up of Highland cow (Bos taurus) showing thick insulating hair covering face. Isle of Lewis, Outer Hebrides, Scotland, UK, April
Close-up of the nose of a Highland cow (Bos taurus) Isle of Lewis, Outer Hebrides, Scotland, UK, April
Mountain hare (Lepus timidus) alert portrait in white winter coat, Monadhliath Mountains, Scotland, UK, February. Non-ex
Adult male Proboscis Monkey {Nasalis larvatus} head portrait, mangrove forest, Bako NP, Sarawak, Borneo
European river otter (Lutra lutra) swimming with head just above surface, river, Dorset, UK, November
Gloucester old spot domestic pig (Sus scrofa domestica) portrait with mouth open and ring in nose, UK
Male Bearded Pig (Sus barbatus) at forest wallow. Bako NP, Sarawak, Borneo, Malaysia
Close up of muzzle and mane of Shetland Pony, Shetland Islands, Scotland, UK
Raccoon (Procyon lotor) approaching camera, Stanley park, Vancouver, British Columbia, Cananda, September
Polar bear (Ursus maritimus) head portrait of adult scenting the wind, Arctic National Wildlife Refuge, Alaska
Male Saiga antelope (Saiga tatarica) running, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009. WWE OUTDOOR EXHIBITION. WWE BOOK
Masked crab (Corystes cassivelaunus) on sandy seabed, Studland Bay, Dorset, UK, May
Wild Boar (Sus scrofa) female smelling air for scent of human, Forest of Dean, Gloucestershire, UK, March
Wild boar (Sus scrofa) female in woodland undergrowth sniffling air for scent of human, Forest of Dean, Gloucestershire, UK, March, Highly commended, Animal Portraits
Adder (Vipera berus) head portrait, Staffordshire, England, UK, April
A large Great hammerhead shark (Sphyrna mokarran), about 4m in length, cruises overhead. Bahamas, Tropical West Atlantic Ocean
Large Tree Shrew (Tupaia tana) Captive, occurs Borneo and Sumatra
Blue Shark (Prionace glauca) near sea surface and divers in background. Santa Maria, Azores, September
Domestic pig, hybrid large white sow in free-range sty, UK, August 2010
Domestic pig, free range sow and piglet sniffing noses, UK, August 2010
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
A domestic pig (Sus scrofa domestica) swimming in the sea. Exuma Cays, Bahamas. Tropical West Atlantic Ocean. This family of pigs live on this beach in the Bahamas and enjoy swimming in the sea
Moai stone statue at dusk on the slopes of the quarry at Rano Raraku volcano, Easter Island Pacific ocean, November 2004
A close up of a blue shark (Prionace glauca) as it investigates the camera beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August
RF- Free range Domestic pig (Sus scrofa domesticus) waiting for feed, UK, August 2010
Leaf nosed twig snake (Langaha madagascariensis) male, Baie de Baly National Park, North west Madagascar
Saiga antelope female {Saiga tatarica}
Young Grey seal (Halichoerus grypus) taking a curious peep out of the water, Island of Coll, Inner Hebrides, Scotland, UK, July
Wild boar (Sus scrofa) female, Forest of Dean, Gloucestershire, UK, March
Grey wolf (Canis lupus) close up, captive
Two female Leaf nosed twig snakes {Langaha madagascariensis} Mantadia NP, East Madagascar
Close-up of Brown bear (Ursus arctos) in lake, Kamchatka, Far east Russia, August
Southern elephant seal (Mirounga leonina) close up of sleeping individual, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska
Close up head portrait of Caracal (Caracal caracal) captive
Head portrait of male Mandrill (Mandrillus sphinx) Captive. Apenheul zoo; the Netherlands
Grey wolf (Canis lupus) head, captive
Highland cow (Bos taurus) close up of head, Scotland, UK
RF- Konik Wild Horse (Equus ferus caballus) low angle close up of head. The Netherlands. July
Close up of nostrils of White horse of the Camargue on wetlands, Camargue, France, January 2009