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Brown hare (Lepus europaeus) adult running in wintery landscape. Derbyshire, UK, January
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Otter (Lutra lutra) female grooming in seaweed, Mull, Scotland, England, UK, September
Mountain hare (Lepus timidus) running in half summer coat, Peak District, UK April
Grass Snake (Natrix natrix) on lily pad, reflected in water. Leicestershire, UK, October
Male Adder (Vipera berus) tasting the air. Cannock Chase, Staffordshire, UK, October
Hazel Dormouse (Muscardinus avellanarius) hibernating amongst leaves and acorns. Captive. Leicestershire, UK, October
Noctule bat (Nyctalus noctula) Captive, UK
Blood-vein moth (Timandra griseata) on fallen leaves, Nottingham, UK, August
RF- Little Owl (Athene noctua) perched on tree stump covered in moss. Worcestershire, England, UK. May 2013
RF - Red Grouse (Lagopus lagopus scotica) among heather. Scotland. August
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Puffin (Fratercula arctica) standing on Sea thrift (Armeria maritima) with wings out stretched with a beak full of sand eels. Close-up portrait. Fair Island, Shetland Islands, Scotland, July
Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January
Gannet (Sula bassana) diving, July UK Shetland
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
Southern stingray (Hypanus americanus) female swimming over a shallow sand bank, under morning sun, North Sound, Grand Cayman, Cayman Islands. British West Indies, Caribbean Sea
Hooded nudibranchs (Melibe leonina) filter feeding beneath bull kelp. Hussar Point, Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
RF - Close up detail of the eye and skin pattern of a Giant hawkfish (Cirrhitus rivulatus). Socorro Island, Pacific Ocean
RF - Lemon shark (Negaprion brevirostris) swimming through sun rays in the late afternoon. Little Bahama Bank, Bahamas. West Atlantic Ocean
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
Emerald tree Boa (Corallus batesii) close up portrait of juvenile whilst hanging on a branch. Tambopata, Madre de Dios, Peru
Toco Toucan (Ramphastos toco) head and beak profile portrait, Brazil
Gemsbok (Oryx gazella) silhouetted at dawn, Kalahari Desert, Botswana
American goldfinch (Carduelis tristis) male in breeding plumage, perched in Eastern redbud flowers in spring, New York, USA May
RF - Sunflowers (Helianthus annuus) in bloom on Norfolk, England, UK. August
RF - Coquerels sifaka (Propithecus coquereli) jumping, Anjajavy Private Reserve, north west Madagascar
Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species
Purple sea pen (Virgularia gustaviana) on sandy sea bed. Rinca, Indonesia
RF - Oak (Quercus robur) acorns in autumn, England, UK, August
RF - Marbled white butterfly (Melanargia galathea) on knapweed, with soft focus leaves, Aosta Valley, Gran Paradiso National Park, Italy
Palm tree branch dipping towards water, Puerto Jimenez, Golfo Dulce, Osa Peninsula, Costa Rica
Forest reindeer, (Rangifer tarandus fennicus) in velvet, Viiksimo, Kuhmo region. Finland, July. Rare subspecies which were nearly extinct in the 19th Century
Fork-leaved sundew (Drosera bipinata) leaves unfurling. Cultivated species, occurs in Australia and New Zealand
Wild Garlic / ramsons (Allium ursinum) leaves emerging in early spring. Peak District National Park, Derbyshire, UK. March
The Northern Lights / Aurora Borealis over a river, Vik, Iceland. February 2014
Panther Chameleon {Furcifer pardalis} close-up of skin, green colouration. Masoala Peninsula National Park, north east Madagascar
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Stars and Milky Way over Durdle Door and the Jurassic Coast, with the lights of Weymouth and Portland beyond, Dorset, England, UK, December 2013
