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A misty morning over a mixed woodland in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011. Highly commended in the Wild Woods Category of the British Wildlife Photography
RF- Head portrait of a European grey wolf (Canis lupus), with a second in the background, captive, Norway, February
Ice stalagmites / ice spikes formation, Lake Baikal, Siberia, Russia, March
Black flat-coated retriever dog splashing in water and retrieving stick (Canis familiaris)
Green anaconda (Eunectes murinus) underwater, flicking tongue, Formoso River, Bonito, Mato Grosso do Sul, Brazil
Two Irish Gypsy cob mares (Equus caballus), one dapple grey - grazing and one piebald - walking on rough pastureland on Hackpen Hill with a Beech clump (Fagus sylvaticus) in the background
Sandstone cliffs at West Bay, Jurassic coast, Bridport, Dorset, UK, May
River Gave de Gavarnie with Cirque du Gavarnie in the distance, Pyrenees National Park, France, June 2013
European common frog (Rana temporaria) on rock by waterfall, near Lac des Gloriettes dam, Pyrenees National Park, France, June
Black veined white butterfly (Aporia crataegi) on grass, Gavarnie-Gedre, Pyrenees National Park, France, June
Geranium Argus butterfly (Plebejus eumedon) feeding on Wood Cranesbill (Geranium sylvaticum) northern Finland, June
Rough seas at Nividic lighthouse during Storm Ruth, Ile d Ouessant, Armorique Regional Park, Iles du Ponant, Finistere, Brittany, France, Iroise Sea. 8th February 2014
Rough seas at Creac h lighthouse during Storm Ruth, Ile d Ouessant, Armorique Regional Park, Iles du Ponant, Finistere, Brittany, France, Iroise Sea. 8th February 2014
Humpback Whale (Megaptera novaeangliae) tail slapping, tail lob, lobtailing. Vancouver Island, British Columbia, Canada, July
Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa
Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Tree sparrows (Passer montanus) perching on a branch in the rain. Perthshire, Scotland, December
Tabby cat, Max, 5 months, with smoke black kitten
Chacma baboon (Papio hamadryas ursinus) portrait of adult male. deHoop Nature reserve. Western Cape, South Africa
Grey heron (Ardea cinerea) adult in breeding plumage, close-up of head and colourful orange beak, backlit, Berlin Zoological Garden, Berlin Zoo, Germany. February
Lion (Panthera leo) male shaking his head after rain, Masai-Mara Game Reserve, Kenya
Starfish Beach, with many starfish in the shallow sea (Asteroidea) Colon Island, Bocas del Toro Archipelago, Bocas del Toro Province, Panama
White-tailed eagle (Haliaeetus albicilla) in flight, Norway, April
Alpaca Herd on Norfolk Farm, winter
Polar bear (Ursus maritimus) portrait, female. Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Lesser flamingo (Phoeniconaias minor) portrait, Lake Bogoria, Rift valley, Kenya, Africa
Portrait of brown and white coated Holland lop-eared domestic rabbit (Oryctolagus cuniculus domesticus), Lorraine, France, July
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Fresh Wych elm (Ulmus glabra) leaves in spring. Dorset, UK May
Lion(Panthera leo) cub playing with its mothers tail, Masai-Mara Game Reserve, Kenya. Vulnerable species
RF -Three colourful Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Blakistons Fish Owl (Ketupa blakistoni) perched, Hokkaido, Japan, February
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Steppe eagle (Aquila nipalensis) in flight, Oman, November
Golden Eagle (Aquila chrysaetos) in flight, snowing, Utajarvi, Finland, February
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
RF- Golden eagle (Aquila chrysaetos) landing in snow, Flatanger, Norway. November
Ash Tree (Fraxinus excelsior) growing in a field. Peak District National Park, Derbyshire, UK. August
RF - Red Deer stags (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Swallowtail butterfly (Papilio machaon) in garden, Norfolk, England, UK, June
RF - Female emperor dragonfly (Anax imperator) laying eggs on garden pond, Broxwater, Cornwall, UK. July
RF - European Badger (Meles meles) foraging at tree trunk in deciduous woodland. Mid Devon, UK. June
Great crested newt (Triturus cristatus) male in breeding condition, amongst Soft hornwort (Ceratophyllum submersum), Herefordshire, England, UK, May. Captive
Red fox (Vulpes vulpes) peering from behind tree with bluebells in foreground, Cheshire, June
Emperor moth (Saturnia pavonia) female wide angle view showing heather moorland habitat. Peak District National Park, UK. April
Red squirrel (Sciurus vulgaris) feeding, Black Isle, Scotland, UK, February
Brown hare (Lepus europaeus) adult running in wintery landscape. Derbyshire, UK, January