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Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
Llama (Lama glama), Los Glaciares National Park, Argentina
The forests of the Modi Khola river valley shrouded in fog. Annapurna Sanctuary, central Nepal, November 2011
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
RF - Red Squirrel (Sciurus vulgaris) on log in snow. Scotland, UK. December
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Welsh Terrier, bitch with puppies aged 8 weeks in snowy scene
Red deer stag (Cervus elaphus) in heavy snow, Cheshire, UK, March
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
Chlorophyte or green alga (Volvox aureus) in pond water. UK. The larger spheres are approximately 0.5mm in diameter. Digital composite image
Bulb tentacle sea anemone (Entacmaea quadricolor) close up detail, Maldives, Indian Ocean
Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February
Twin Birch (Betula) trees highlighted snow. Senja, Norway, February
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Female Scottish wildcat (Felis silvestris grampia) portrait, in winter coat, semi habituated living wild in native oakwood, Lochaber, Scotland, April
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Kilchurn Castle on Loch Awe, Argyll and Bute, Strathclyde, Scotland, UK, April 2011
Adult Adder (Vipera berus) basking on pebble beach. Abbotsbury, Dorset, UK, March
Storm clouds building up off coasts of Norfolk, Weybourne Cliffs, Norfolk, UK, November
Detail of frost on Western Red Cedar (Thuja plicata). Vancouver Island, Canada, March
Colourful water-smoothed pebbles on shingle beach. Normandy, France, October 2010
Wild Horses / mustangs, two stallions rearing up fighting, Pryor Mountains, Montana, USA, June
Pink abalone (Haliotis corrugata) shell detail, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, January
Suffolk Punch heavy horse in field resting head anothers back, UK, September
Norfolk terrier portrait, lying in autumn foliage
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013
Beech (Fagus sylvatica) woodland in spring, at dawn, Peak District National Park, Cheshire, UK, May
Hydromedusa / Trachymedusa (Crossota sp) showing tentacles, from approx 2700m, mid Atlantic ridge, June 2010
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
King cobra (Ophiophagus hannah) in strike pose, Malaysia
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Devils Marbles, Devils Marbles Conservation Reserve, Northern Territory, Australia, June 2010
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February