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European Cave Spider (Meta menardi) in limestone cave. Plitvice Lakes National Park, Croatia. January
Violet bramble rust (Phragmidium violaceum) causing red colouration on Bramble leaf (Rubus fruticosus aggregate). Peak District National Park, Derbyshire, UK. October
Spider hunting wasp (Pompilidae) Maliau Basin, Sabah, Borneo
Stormy light over fields near Longnor, Peak District National Park, Staffordshire, May 2014
Buff-tailed Bumblebee (Bombus terrestris) worker feeding on Ox-eye Daisy (Leucanthemum vulgare). Peak District National Park, Derbyshire, UK. July
Alpine meadow in flower including Scabious flowers. Austrian Alps at 1700 metres altitude, North Tyrol, Austria, June
Lobster pots on harbourside at night, Tobermory harbour, Isle of Mull, Scotland
Indian moon / Indian luna Moth (Actias selene) close up of defensive spines on caterpillar. Captive
RF - Red Squirrel (Sciurus vulgaris) adult in summer coat leaping onto fallen log. Scotland, UK. September
Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015
Raft Spider (Dolomedes fimbriatus) female on heathland pool. Note wing of predated damselfly floating on waters surface under the spider. Dorset, UK. August
Water vole (Arvicola amphibius) feeding at edge of water, Kent, UK, December
Sand lizard (Lacerta agilis) male basking. Dorset, England, UK, May
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Badger (Meles meles) at entrance to sett, Derbyshire, England, UK, June. Highly commended in the Habitat category of the BWPA competition 2014
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Red Squirrel (Sciurus vulgaris) adult sitting and feeding in snowfall. Cairngorms National Park, Scotland, UK, March
Water vole (Arvicola amphibius) standing on hind legs sniffing blackberry, Kent, UK, September
Bank vole (Clethrionomys glareolus) feeding on blackberries, Worcestershire, England, UK, August. Meetyourneighbours.net project
Red Fox (Vulpes vulpes) portrait in an urban area. Glasgow, Scotland. May
Mountain Hare (Lepus timidus) on moorland with white winter coat, Kinder Scout, Peak District National Park, Derbyshire, UK, February
Sanderling (Calidris alba) running along the tide-line, reflected in the mud, Norfolk, England, UK. December
Robin (Erithacus rubecula) singing, Norfolk, England, UK, February
Eyes of Margined sole fish (Brachirus heterolapis) on its camouflaged body. Saonek Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean. Dampier Strait
Reticulate stargazer (Uranoscopus sp. ) emerging from the sandReticulate stargazer (Uranoscopus sp.) emerging from the sand. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean
Red Irish lord (Hemilepidotus hemilepidotus) hides on the sebed, with Quillback rockfish (Sebastes maliger) and copper rockfish (Sebastes caurinus) behind in the bull kelp forest
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF - Split level photo of a Whale shark (Rhincodon typus) feeding at the surface. Isla Mujeres, Yucatan Mexico. Caribbean Sea
RF - Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF - Green sea turtle (Chelonia mydas) beneath clouds. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
Blackbird (Turdus merula) male foraging on snow covered ground, during a blizzard. Derbyshire, UK. January (non-ex)
Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
RF - Early Purple Orchid (Orchis mascula) in flower, Broxwater, Cornwall, UK. May 2016
Lavender (Lavandula angustifolia) fields, Valensole Plateau, Alpes Haute Provence, France, June
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
Great white egret (Ardea alba) silhouetted at sunset, Lake Csaj, Pusztaszer, Hungary, February. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Ruppells weaver (Ploceus galbula) male building nest, Oman, May
RF - Marbled White butterfly (Melanargia galathea) resting on common spotted orchid, Dunsdon Nature Reserve, Devon, UK. July
RF- Daffodils (Narcissus) in flower photographed using soft focus technique
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
