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Brown bear (Ursus arctos) and Grey wolf (Canis lupus) together in wetlands, Kuhmo, Finland, July
Five Cavalier King Charles Spaniel sitting, black-and-tan, tricolour, blenheim and ruby coated, sitting in autumn leaves
Silver Studded Blue (Plebejus argus), Gran Paradiso National Park, Aosta Valley, Pennine Alps, Italy. July
Rough seas at d'Ar-Men lighthouse during Storm Ruth, Ile de Sein, Armorique Regional Park. Iles du Ponant, Finistere, Brittany, France, Iroise Sea. 8th February 2014
Gemsbok (Oryx gazella) in sand dunes, Namibia. Finalist in the Mammals Category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
View of Salcombe and harbour from Snapeaes Point in the early morning light. Salcombe, South Devon, UK, September 2010
Portrait of German Shorthair Pointer in woodland, Illinois, USA
RF - Dapple grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
Hot air balloon over the Temples of Bagan at dawn, Myanmar, November 2012
Northern / Hen Harrier (Circus cyaneus) and Kestrel (Falco tinnunculus) below, fighting in flight. Vosges, France, February
Harpy Eagle (Harpia harpyja) portrait. Gamboa, Soberania National Park, Panama
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Female Anglerfish fanfin seadevil (Caulophryne jordani) - Deep sea species from Atlantic Ocean, found at 700-3000m
Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Chilean flamingos (Phoenicopterus chilensis) in flight over mountain peaks with glacier in the distance, Torres Del Paine National Park, Chile
Flemish giant rabbit with ears erect
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Fingals Cave, showing basalt columns, Isle of Staffa, Inner Hebrides, Scotland, UK, June 2010
Leatherback Turtle (Dermochelys coriacea) hatchling crossing a beach to get to the sea. Cayenne, French Guiana, July
RF - Grey Andalusian mare running in snow, Berthoud, Colorado, USA. January
Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA
Red deer (Cervus elaphus) stag in the snow, Scotland, March
RF - Lions (Panthera leo) - two brothers patrolling territorial boundary. Short grass plains on the border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Wild Siberian / Amur tiger (Panthera tigris altaica) in woodland, near Perekatnaj river, Lazovskyi region, Primorskyi krai, Far East Russia, February. Endangered species
Volcanic eruption, Eyjafjallajokull, near the Myrdalsjokull glacier, South Iceland, april 2010
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Orangutan baby (Pongo pygmaeus) head portrait of baby, Semengoh Nature reserve, Sarawak, Borneo, Malaysia. Endangered
African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012
Red fox (Vulpes vulpes) cubs playing, Vosges, France, May
Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK. January
King penguins (Aptenodytes patagonicus) crossing water to reach breeding site, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Portrait of a young Little Owl (Athene noctua)
Wandering albatross {Diomedea exulans} flying over open ocean, South Atlantic
Buachaille Etive Beag reflected in Lochan na Fola after snowfall, early morning light, Glencoe, Scotland, UK. March 2017
Crowns Engine house, abandoned mining buildings, on coastasl cliffs at Botallack head, Cornwall, UK. February 2017
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Portrait of English Springer Spaniel (field type). Elkhorn, Wisconsin, USA, January
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Malaysian Orchid Mantis (Hymenopus coronatus) pink colour morph, camouflaged on an orchid. Captive, from Malaysia
Cavalier King Charles Spaniel puppies aged 7 weeks, with tricolour and blenheim colouration, on hay bale in straw
Japanese macaque (Macaca fuscata) sleeping at hot spring in Jigokudani, Yaenkoen, Nagano, Japan, February
Blacktail Prairie Dog (Cynomys ludovicianus) engaging in Jump-yip behavior - A strong arch of the back or " jump" followed by a shrill " yip" - Thought to occur when a predator
Tortoiseshell kitten, standing, against white background
Malaysian Orchid Mantis (Hymenopus coronatus) white colour morph camouflaged on an orchid. Captive. Originating from Malaysia
RF - Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April