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Edible chanterelle fungus (Cantharellus cibarius) showing spore dispersal pattern over 24 hours on black card
False death cap fungus (Amanita citrina var. alba) showing spore dispersal pattern over 22 hours on black card
A cascade of waterfalls falling over a cliff in woodlands. Plitvice National Park, Croatia, May 2010
A view of a forested inlet lit by low sunlight. Aggro Beach, west coast of Vancouver Island, Canada, September 2010
Cosmos (Cosmos bipinnatus), Bachelor buttons / Cornflowers (Centaurea cyanus), and Common poppies (Papaver rhoeas) in a small meadow. Nevada, USA, July
Conifer trees laden with snow in winter landscape, Kuusamo, northern Finland, February 2009
Beginnings of the Northern lights in night sky in winter with conifer trees laden with snow, Kuusamo, northern Finland, February 2009
Common raven (Corvus corax) perched in snow beside birch tree, Finland, March
Poets Daffodil (Narcissus poeticus). Montseny Nature Reserve, Barcelona province, Catalonia, Spain, May
Iceberg against a blue sky. Ilulisat, West Greenland, summer
Silver Birch (Betula verrucosa) reflected in Loch Beinn a Mheadhoin. Glen Affric National Nature Reserve, Scotland, December
Tree covered with rime ice standing in snow-covered field, Ithaca, New York, USA. January 2010
Fragrant water lilies (Nymphaea odorata) in Lake Skadar, Montenegro, May 2008
Alpine meadows with Caucasian scabious (Scabiosa caucasica) in Bezengi gorge, July, 3000m. W-C Caucasus, Kabardino-Balkarskiy nature reserve, Russia
View up through Palm trees in swamp forest, Borneo, Sarawak, Malaysia
Vipers bugloss (Echium vulgare) flowering in an olive grove. Sierra de Andujar Natural Park, Jaen, Andalusia, Spain, April 2010
Alder trees (Alnus sp) reflected in standing water in nature reserve, Berlin, Germany, May 2009
Beech trees (Fagus sylvatica) in West Woods, Wiltshire, UK, November 2010
Lenga forest (Nothofagus pumilio) in Ascensio river valley on the W Trek, Torres del Paine National Park, Patagonia, Chile, January 2006
Tsemo Gompa, seen from the road between Leh and Kardung La, Ladakh, India, June 2010
Aerial view of Dyrholaey and Reynisdrangar, near Vik, in South Iceland, April 2010
Solar eclipse 63% on 4 January 2011, seen from Barcelona, Spain
Mature Beech tree (Fagus sylvatica) with leaves in autumn colours, Mark Ash Wood, New Forest, England, UK, October
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
Panoramic view, Little Razorback Island, Dellbridge Islands, Antarctica, October 2008, Digitally stitched image
Ivy (Hedera helix) leaves with Star moss (Polytrichum) and Feather moss (Thuidium tamariscum) Kingcombe, Dorset, UK March 2010
Clouds during the rainy season in the Kalahari Desert, Botswana, March 2009
Catesbys snail-eating snake (Dipsas catesbyi) on Heliconia plant, Pacaya Samiria National Reserve, Amazon Basin, Loreto, Peru
Golden eagle (Aquila chrysaetos) in flight over snow, Alps, Austria, controlled conditions
Panther Chameleon {Furcifer pardalis} walking along branch on coast, Masoala Peninsula National Park, north east Madagascar
Tropical rainforest meeting the sea. Masoala Peninsula National Park, north east Madagascar, October 2009
Boabab trees {Adansonia grandidieri} in evening light. Morondava, Madagascar. Digital Composite Panorama
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Eye and spiracle detail of a Bluespotted / Ribbontail Stingray (Taeniura lymma) Sipadan Island, Sabah, Borneo, Malaysia. May
California sea lion pup (Zalophus californianus) portrait in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Atlantic grey seal (Halichoerus grypus) swimming beneath the surface, Lundy Island, Devon, England, UK. July 2010. Highly commended, Coast and Marine category
Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean
Black browed albatross (Thalassarche melanophrys) flying in 60 knot wind at Cape Horn, Chile
Lemon shark (Negaprion brevirostris) in shallow water with reflection at the surface. Little Bahama Bank, Bahamas
Caribbean reef shark (Carcharhinus perezi) swims over a coral reef. Grand Bahama Island, Bahamas. Tropical West Atlantic Ocean
Bottlenose dolphins (Tursiops truncatus) in shallow water over a sand bank. Sandy Ridge, Little Bahama Bank, Bahamas. Tropical West Atlantic
Polar bear (Ursus maritimus) footprints in the snow along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska. October 2009
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska
Common / Eurasian cranes (Grus grus) small flock of juveniles taking flight over the frozen, snow covered pastureland on a foggy winter morning
Red Kite (Milvus milvus) diving towards prey whilst in flight, Wales, UK, March
Crested Newt (Triturus cristatus carnifex) underwater view of female swimming on surface, Traversari lake, Campigna National Park, Emilia Romagna and Tuscany regions, Italy
Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive
As soon as it entered the water the young Olive ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the coast
