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Honey garlic (Nectaroscordium siculum) with Bumblebee in flight. Naturalised in the Avon Gorge, Bristol, UK, June
Common snowdrops (Galanthis nivalis) growing in the Rococo Gardens, Painswick, Gloucestershire, Engalnd, UK. February
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
RF - Male African lion (Panthera leo) scratching tree at Big Marsh, Nogorongoro Conservation Area / Serengeti National Park, Tanzania. March
RF - Andean condor (Vultur gryphus) flying over Torres del Paine Massif. Torres del Paine National Park, Chilean Patagonia, Chile. March
RF- Verreauxs sifaka lemur (Propithecus verreauxi) hanging from branch while feeding in canopy. Berenty Private Reserve, southern Madagascar. Endangered species
RF- Stump tailed chameleon (Brookesia superciliaris) portrait, Andasibe-Mantadia National Park, Eastern Madagascar
RF- Morpho butterfly displaying on leaf (Morpho achilles). Amazonia, Ecuador
African jackass penguins (Spheniscus demersus) three heading to sea in the early morning to forage for food, Cape Town, South Africa, Endangered IUCN
RF - Split level photo of a coral reef with hard corals (Acropora sp. Millepora sp. and Pocillopora sp.) and the shore with palm trees, at sunset. Red Sea
RF - Red fox (Vulpes vulpes) foraging in snow covered valley. Hayden Valley, Yellowstone National Park, USA. February 2019. (This image may be licensed either as rights managed or royalty free.)
RF - King penguin (Aptenodytes patagonicus), four returning to sea. St Andrews Bay, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
Steam rising from Grand Prismatic thermal pool in snow covered landscape. Midway Geyser Basin, Yellowstone National Park, USA. February 2019
Chinese goral (Naemorhedus griseus) in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Snow partridge (Lerwa lerwa) flock flying past snowy rockface, in evening light. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Honey bee (Apis mellifera) covered in pollen emerging from nectaring in Crocus (Crocus sp). On nature reserve, Surrey, England, UK. March
Common bee fly (Bombylius major) nectaring on Primrose (Primula vulgaris). This insect is the main pollinator of Primrose. In nature reserve, Surrey, England, UK, April
Red mucronata (Balanophora harlandii), a parasite on woody roots with flowers resembling fungi. Wild Elephant Valley, Xishuangbanna, Yunnan, China
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
China nawab (Polyura narcaeus) butterfly drinking at thermal seep. Gaoligongshan National Nature Reserve, Yunnan Province, China
China glory flower (Clerodendrum bungei) fruits above red calyxes. Eaten and spread by birds. Tangjiahe National Nature Reserve, Sichuan Province, China
Flowers and leaves on forest floor. Xishuangbanna National Nature Reserve, Yunnan, China
Sancha River flowing through tropical rainforest in dry season. Wild Elephant Valley, Xishuangbanna National Nature Reserve, Yunnan Province, China. April 2007
Pyramid of sea ice catching the first rays of the returning sun in the high Arctic, Svalbard, Spitsbergen, Norway, February 2009. Taken on location for the BBC series, Frozen Planet
Dead leaf / Orange oak leaf (Kallima inachus) butterflies feeding on sap on tree trunk. Mimic leaves with wings closed. Wild Elephant Valley, Xishuangbanna, Yunnan Province, China
Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September
Barn owl (Tyto alba) perched on fence post at dusk, with industrial estates of Teeside in the background. Durham, UK. February
Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka
Forsters hawkfish (Paracirrhites forsteri) on a coral reef. Jackson Reef, Tiran, Sinai, Egypt. Strait of Tiran, Gulf of Aqaba, Egypt
Hawksbill turtle (Eretmochelys imbricata) unusually feeding on hard coral polyps. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Green turtle (Chelonia mydas) descending after breathing at the surface. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Three Pink anemonefish (Amphiprion perideraion) living in a purple skirted magnificent sea anemone (Heteractis magnifica) on a coral reef, below a small island
Tawny nurse sharks (Nebris ferrungineus) at night. Vavuu Atoll, Maldives. Indian Ocean
Midnight snapper (Macolor macularis) shelters in a cavern on a coral reef with white soft corals (Scleronephthya sp.) and solderfish. Vavuu Atoll, Maldives. Indian Ocean
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Reflections in autumn leaves in lake, Acadia National Park, Maine, USA, October
Waterfalls flowing into Hvita River at Hraunfossar, Iceland, September 2013
RF- Moorland view from Littaford Tor, near Two Bridges, Dartmoor National Park, Devon, England, UK, October 2012
RF- Red Admiral butterfly (Vanessa atalanta) perched on Bigtooth Maple (Acer grandidentatum). Lost Maples State Park, Hill Country, Central Texas, USA. November
A high view of mist shrouding an autumnal valley. Cadi Natural Park, Catalonia, Barcelona province, Pyrenees, Spain, October 2005
RF- Northern Cardinal (Cardinalis cardinalis) immature male perched on Texas Red Oak (Quercus Texana). New Braunfels, San Antonio, Hill Country, Central Texas, USA
RF- Mountain bearberry (Arctous alpinus) covered in frost, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden. September
RF- American robin (Turdus migratorius) female feeding in Black hawthorn (Crataegus douglasii). Grand Teton National Park, Wyoming, USA
RF- Reindeer (Rangifer tarandus) feeding. Denali National Park, Alaska, USA
Snares Crested penguin (Eudyptes robustus) bathing in freshwater pool, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Potrait of a Cuckoo wrasse (Labrus bimaculatus) Loch Linnhe, west coast of Scotland, UK, September