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Aerial view of low cloud over the River Spey meandering through Insh marshes, Cairngorms National Park, Scotland, UK. May, 2017
Whooper swans (Cygnus cygnus) on snow covered Loch Insh in winter, Cairngorms National Park, Scotland, UK. January
Atlantic salmon (Salmo salar) leaping up waterfall, Cairngorms National Park, Scotland, UK. October
Male Black grouse (Tetrao tetrix) performing courtship display at lek, Cairngorms National Park, Scotland, UK. April
Alder (Alnus glutinosa) tree silhouetted by a display of Aurora borealis in the night sky, Cairngorms National Park, Scotland, UK. February
Two Coal tits (Periparus ater) perched on a branch in falling snow, Cairngorms National Park, Scotland, UK. December
White-tailed deer (Odocoileus virginianus) doe with fawn in woodland, Acadia National Park, Maine, USA. July
American Purple gallinule (Porphyrio martinica) balancing on a lily pad, Everglades National Park, Florida, USA
Great blue heron (Ardea herodias) taking flight, Acadia National Park, Maine, USA. June
Aurora borealis in the evening sky over Ben Ledi from Buchlyvie, The Trossachs National Park, Stirling, Scotland. September, 2020
Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp)Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp.), Kubah National Park, Sarawak, Borneo
Wolf (Canis lupus) pack huddling together in snow-covered forest, Sumava National Park, Bohemian Forest, Czech Republic. Captive
Two Great white egret (Ardea alba) in winter, Kiskunsag National Park, Hungary. December
Grey heron (Ardea cinerea) on lake at sunset, Lake Csaj, Kiskunsagi National Park, Hungary. January
Close up of a Forest elephant (Loxodonta africana cyclotis) hidden in foliage, Loango National Park, Gabon. December
Lowland gorilla (Gorilla gorilla) mother and young in forest, Loango National Park, Gabon. January
Leopard (Panthera pardus), Masai Mara National Park, Kenya. July
Asiatic lion (Panthera leo persica) cub, Gir Forest National Park, Gujarat, India. May
Greater one-horned rhinoceros (Rhinoceros unicornis) close up, Kaziranga National Park, India. January
RF - Great Indian Hornbill (Buceros bicornis) in forest canopy. Kaziranga National Park, Assam, India
RF- Portrait of male Parsons Chameleon (Chamaeleo parsonii), Ranomafana National Park, South eastern Madagascar
RF- Adult male Serval (Felis serval) stalking prey in long grass. Serengeti National Park, Tanzania
Dominant male Chimpanzee (Pan troglodytes troglodytes) walking in the mangrove, with female carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Female Chimpanzee (Pan troglodytes troglodytes) climbing in trees carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Russel Falls, Mt Field National Park, Tasmania, Australia. April, 2016
Female Yellow-footed rock wallaby (Petrogale xanthopus) fending off advances from male (right), Idalia National Park, Queensland, Australia
Infant Chimpanzee (Pan troglodytes troglodytes) climbing in tree, Conkouati-Douli National Park, Republic of Congo, Africa
Aerial view of mangrove forest, Conkouati-Douli National Park, Republic of Congo, Africa. October, 2021
Dominant male Chimpanzee (Pan troglodytes troglodytes) in the mangrove, with female and infant vocalising, Conkouati-Douli National Park, Republic of Congo, Africa
Male Saiga antelope (Saiga tatarica) rutting in winter, The Black Lands National Park, Kalmykia, Russia
Hooded grebes (Podiceps gallardoi) pair dancing in breeding display, on a lake above the Buenos Aires plateau, Patagonia National Park, Santa Cruz, Argentina. Cropped
Hooded grebes (Podiceps gallardoi) pair on lake, with mountains in Chile in background, Buenos Aires plateau, Patagonia National Park, Santa Cruz, Argentina. January 2018
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
Smoke billowing from Mount Bromo volcano. View from the summit of Mount Batok. Bromo Tengger Semeru National Park, Java, Indonesia, July, 2013
Eastern bentwing / Eastern long fingered bat (Miniopterus orianae oceanensis) hibernating cluster in cave, one bat with bat fly (Hippoboscoidea) Girraween National Park, Queensland, Australia
Eastern long fingered bat (Miniopterus orianae oceanensis) roosting in a cluster on roof of a cave during day, a few ectoparasitic batflies (Hippoboscoidea) are visible, Girraween National Park
Alpine ibex (Capra ibex) adult males fighting on a steep mountain side, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Escuain Canyon, Autumm, Ordesa National Park, Pyrenees, Aragon, Spain
Beech (Fagus sylvatica), large tree amongst Bracken (Pteridium aquilinum). Bolderwood, New Forest National Park, Hampshire, England, UK. October 2020
Autumn colours in Mountain birch, (Betula pubescens var. tortuosa), Abisko National Park, Norrbotten, Lapland, Sweden September 2020
Reflections in the salt marsh at Penouille at dawn, Forillon National Park, Gaspesie, Quebec, Canada. October 2019
Curbar Edge, Peak District National Park, Derbyshire, England, UK. April 2018
Badwater, the lowest point in the the USA and the hottest place on earth, Death Valley National Park, California, USA
Mixed decidious forest, autumm, Ordesa National Park, Aragon, Pyrenees, Spain
Beautiful hook-tip moth (Laspeyria flexula). Peak District National Park, Derbyshire, UK. June. Focus stacked image
Close-up of scales on underside of Red admiral butterfly wing (Vanessa atalanta). Peak District National Park, Derbyshire, UK. August. Focus stacked image
Tigress drinking water (Panthera tigris tigris) hidden in long grass, Tadoba National Park and Tiger Reserve, Maharashtra, India. March
Bengal tiger (Panthera tigris) cooling down in waterhole, portrait. Ranthambore National Park, India