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Oystercatchers (Haematopus ostralegus) arriving to roost at Snettisham on the Wash. Norfolk, January
Flock of Knot (Calidris canuta) over sea with wind turbine. Liverpool Bay, UK, December
Five images of schooling Raccoon Butterflyfish (Chaetodon lunula) digitally combined for this panorama photograph. Lanai, Hawaii
Schooling Black Striped Salema / Snapper (Xenocys jessiae) and Yellowtail Surgeonfish (Prionurus laticlavius). Galapagos Islands, Equador. Three images were stitched to create this panorama
Flock of Oystercatchers (Haematopus ostralegus) and Knot (Calidris canuta) in flight and on tidal shallows. The Wash Estuary, Norfolk, September
Gannet (Morus bassanus) breeding colony, long exposure showing flight trails. Hermaness National Nature Reserve, Shetland, Scotland, July
Flock of Knot (Calidris canuta) taking flight. Snettisham, UK, January
Large flock of Barnacle geese (Branta leucopsis) taking off from grazing marshes, Caerlaverock WWT, Dumfries and Galloway, Scotland, UK, December
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November