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Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
School of Blackfin barracuda (Sphyraena qenie), Kimbe Bay, West New Britain, Papua New Guinea
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Brown-lipped snail (Cepaea nemoralis) on fungus, reproduced in a bokeh affect, Sheffield, England, UK. November. Commended in the Natures Studio Category of GDT competition 2014
Herd of Tibetan Wild Asses / Kiang (Equus kiang) ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India, October 2012
Nile crocodile (Crocodylus niloticus) making its way through sand to the Rufiji River, Selous Game Reserve, Tanzania
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
Male Bengal tiger (Panthera tigris tigris) known as Sundar (B2) patrolling territory, Bandhavgarh National Park, Madhya Pradesh, India. Endangered species
RF- Bottlenose dolphin (Tursiops truncatus) swimming in a spiral movement, Sandy Ridge, Little Bahama Bank. Bahamas
Gentoo penguins (Pygoscelis papua) returning over sea ice to breeding colony, Signy Island, South Orkneys, Antarctica, December
Arctic fox (Alopex lagopus) in winter coat, running across snow, Svalbard, Norway. April
Polar bear (Ursus maritimus) female with cub walking over snow with mountains in background, Svalbard, Norway. April
Polar bear (Ursus maritimus) walking across ice and snow in evening sun. Svalbard, Norway. April
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
A shoal of Golden grey mullet (Liza aurata) swim in the surf breaking on the shore of El Hierro Island. Canary Islands, Spain. East Atlantic Ocean
Two Caracal (Caracal caracal) cubs, aged 9 weeks, walking over rocks, Spain. Captive, occurs in Africa and Asia
Leopard (Panthera pardus) female, walking across grassland in morning sunlight, Kenya
Eastern rockhopper penguin (Eudyptes chrysocome filholi) mid-fall after jumping off rocky ledge to get to sea, surrounded by Bull kelp (Durvillaea antarctica), Auckland Island, Auckland Island
Brown bear (Ursus arctos) female with two cubs walking through misty, woodland meadow, Finland. June
Arctic fox (Alopex lagopus) in winter coat, walking across snow, Svalbard, Norway. April
Polar bear (Ursus maritimus) female with cub, walking over snow with mountains in background, Svalbard, Norway. April
Giant anteater (Myrmecophaga tridactyla) walking, Pantanal, Mato Grosso do Sul, Brazil
Iberian ibex (Capra pyrenaica) kid, aged two months, making its way down cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), adult female, walking along coastal cliff. Mediterranean coast, Andalusia, Spain. November
African elephant (Loxodonta africana), female and calf, crossing plain at sunset. Amboseli National Park, Kenya. July
Masai giraffe (Giraffa camelopardalis tippelskirchi) running across plain. Amboseli National Park, Kenya. July. Digitally enhanced
Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp)Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp.) being followed by a California sea lion (Zalophus californianus), Espiritu Santo Island
Water rail (Rallus aquaticus) wading through water, Dumfries and Galloway, Scotland, UK
Sand cat (Felis margarita) walking over sandy ground. Captive, occurs in North Africa, the Arabian Peninsula, Pakistan and the Middle East
Female Olive ridley turtles (Lepidochelys olivacea) coming ashore to nest while others leaves, at sunrise, during massive arribada. Playa Escobilla Sanctuary, Oaxaca, Mexico. Pacific Ocean
Adult American oystercatcher (Haematopus palliatus) walking along beach, with chick running and flapping its wings. Long Island, New York, USA. June
Gemsbok (Oryx gazella) female and calf walking past sand dune, Sossusvlei UNESCO World Heritage Site, Namib desert, Namib-Naukluft National Park, Namibia
Herd of wild Onaqui horses (Equus ferus) trotting in dust. Onaqui Mountain Herd Management Area, Utah, USA. September
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Amur leopard (Panthera pardus orientalis) walking past snow-covered trees on mountain slope, Land of the Leopard National Park, Russian Far East. Critically endangered. Taken with remote camera