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Female Anglerfish fanfin seadevil (Caulophryne jordani) - Deep sea species from Atlantic Ocean, found at 700-3000m
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Serotine bat (Eptesicus serotinus) showing teeth, Captive, UK
Turbot (Scophthalmus maximus) detail of the eyes and mouth, United Kingdom, September
Black rhinoceros {Diceros bicornis} charging with mouth open, Etosha national park, Namibia
RF- Jaguar head profle portrait (Panthera onca) head portrait captive. Pantanal, Brazil
Common raven (Corvus corax) calling, beak open, The Burren, County Clare, Ireland, June 2009
Monkfish / Allmouth (Lophius piscotorius) close-up of face, Lofoten, Norway, November 2008
Basking shark (Cetorhinus maximus) feeding at the surface on plankton, Cairns of Coll, Inner Hebrides, Scotland, UK, June
Atlantic wolf fish (Anarhichas lupus) in a rock crevice, St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, August 2011
European Grey Wolf male yawning {Canis lupus} captive, Transsylvania, Romania
Basking shark (Cetorhinus maximus) feeding on plankton in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June. 2020VISION Book Plate
Leopard (Panthera pardus) female yawning, Samburu game reserve, Kenya
Blue whale underwater close-up of head and mouth, Mexico
Viperfish (Chauliodus sloani) - deep sea specimen from Portugal
Wild goats (Capra hircus) resting on coastal promontory, one yawning, Great Orme, Llandudno, Gwynedd, North Wales, UK, August
Japanese macaque / Snow monkey {Macaca fuscata} mother grooming four-day-old newborn baby, Jigokudani, Nagano, Japan
Great white shark (Carcharodon carcharias) underwater, Guadalupe Island, Mexico (North Pacific)
Head portrait of Sumatran tiger (Panthera tigris sumatrae) with mouth wide open in a yawn, captive
Jaguar head profile portrait {Panthera onca} captive, Pantanal, Brazil
Japanese Macaque (Macaca fuscata) male yawning, Jigokudani Japan, January
Sloth Bear (Melursus ursinus) female with large cubs (on right) trying to fight off rogue male. Karnataka, India, March
Alligator snapping turtle (Macroclemys temmincki) fishing by luring a fish with the vermiform, worm-like, tongue, USA, captive
Mandrill (Mandrillus sphinx) male showing aggression, captive
Close up of mouth sucker of Japanese / Arctic Lamprey {Lethenteron japonicum} Japan
Atlantic wolffish {Anarrhichus lupus} Sognefjord, Norway
Basking shark (Cetorhinus maximus) feeding just below the surface, Mull, Scotland, June 2009
Greater black backed gull (Larus marinus) swallowing large fish, North Atlantic, Flatanger, Nord-Trndelag, Norway, August 2008
Basking shark (Cetorhinus maximus) off the Island of Mull (Coll and Tiree Islands area) Scotland, June 2009 WWE BOOK
Anglerfish (Lophius piscatorius) portrait, Saltstraumen, Bod, Norway, October 2008. WWE INDOOR EXHIBITION
Grey seal (Halichoerus grypus) swimming amongst kelp and playing, Farne Islands, Northumberland, England, UK, July
Velvet Swimming Crab (Necora puber / Liocarcinus puber), St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, October, 2011
Red deer (Cervus elaphus) stag bellowing during autumn rut, Richmond Park, London, England, UK, October
Lion (Panthera leo) male smelling, flehmen response, Masai-Mara Game Reserve, Kenya. Vulnerable species
Grey seal (Halichoerus grypus) laughing, Helgoland, Germany
Spotted hyaena laughing {Crocuta crocuta} Kenya
Old male Grey seal (Halichoerus grypus) with mouth wide open showing worn teeth, Lundy Island, Bristol Channel, England, UK, May. 2020VISION Book Plate
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Orang utan (Pongo pygmaeus) baby hanging from trees, and chewing on fingers, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Lemon shark (Negaprion brevirostris) in shallow water at the surface, split level. Little Bahama Bank, Bahamas. West Atlantic Ocean
Great white shark (Carcharodon carcharias) portrait, Guadalupe Island, Mexico, Pacific Ocean
Head portrait of Sumatran tiger (Panthera tigris sumatrae) with face half cast in shadow, with aggressive expression, captive
Fangtooth (Anoplogaster cornuta) portrait, Mid-Atlantic Ridge, North Atlantic Ocean
Maine Coon female cat, Serafin, in fierce defensive posture
Dalmation yawning portrait
Blue Roan Wild Stallion yawning {Equus caballus} Pryor Mountains, Montana, USA
Arctic fox yawning {Vulpes lagopus} Norway captive