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A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Atlantic salmon (Salmo salar) leaping up waterfall to reach spawning grounds upstream. River Endrick, Loch Lomond and The Trossachs National Park, Scotland, UK. July
Adelie penguin (Pygoscelis adeliae) walking on ice, near Davis Station, Prydz Bay, East Antarctica, November
Low angle view of Demoiselle crane (Anthropoides virgo) flying and feeding during winter migration. Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) low angle view of birds flying and landing near the chugga ghar (bird feeding enclosure ) in Khichan Village, durin their annual migration
Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Portrait of Male Sockeye salmon (Oncorhynchus nerka) showing teeth. Adams River, British Columbia, Canada, October
White stork (Ciconia ciconia) in tree, migrating towards Europe, Masai-Mara Game Reserve, Kenya
Aerial view of migrating Wildebeest (Connochaetes taurinus) on the border of Ngorongoro Conservation Area and Serengeti National Park, Tanzania, Africa
Common cranes (Grus grus) flock on migration at sunset, Montier en Der, Aube, France, November
Grey whale (Eschrichtius robustus) pod migrating north, aerial view. Baja California, Mexico. February
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Gray whale (Eschrichtius robustus) aerial view of blowing during whale migration, Coast of California, USA, Eastern Pacific Ocean
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December
Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya
Grants zebra (Equus burchelli boehmi) in a migration herd of wildebeest (Connochaetes taurinus) Masai-Mara Game Reserve, Kenya
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
