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A shoal of Golden grey mullet (Liza aurata) swim in the surf breaking on the shore of El Hierro Island. Canary Islands, Spain. East Atlantic Ocean
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
Oriental sweetlips (Plectorhinchus vittatus), Maldives, Indian Ocean
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Demoiselle cranes (Anthropoides virgo) flock, India
Brown pelicans (Pelecanus occidentalis) on water, split level view, Galapagos
Lioness (Panthera leo) and Thomsons gazelles (Eudorcas thomsonii) silhouetted at sunset, Masai-Mara game reserve, Kenya
Great egret (Ardea alba) feeding on fish, Pusztaszer, Hungary, June
Group of male Common eiders (Somateria mollissima) in flight, Iceland, June
Group of Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Chapada dos Guimaraes, Brazil
Common Teal (Anas crecca) female with ducklings, Finland June
Lion (Panthera leo) cubs sitting on termite mound, Masai-Mara, Kenya. Vulnerable species
Chihuahuas, mixture of longhaired and short-haired sitting in basket
Lesser flamingos (Phoeiconaias minor) feeding in still lake, Lake Nakuru, Rift valley, Kenya, Africa
Kamchatka Brown Bear (Ursus arctos beringianus) on the Pacific coast of Kamchatka. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Herons at night on Lake Csaj, Kiskunsag National Park, Hungary. Winner of the Birds category, Wildlife Photographer of the Year (WPOY) Competition 2014
Long Horn cattle in meadow, Norfolk, UK July
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
White terns (Gygis alba) flock in flight overhead, Christmas Island / Kiritimati, Pacific Ocean, July
Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May
African moon moth (Argema mimosae) digital composite sequence, showing wings expanding after emerging from cocoon, photographed against a white background, orginating from Africa. Captive
Jersey tiger moth (Euplagia quadripunctaria) large numbers flying to and resting on Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011
Common dolphins ( Delphinus delphis) swimming below the surface, Atlantic Ocean, Portugal, October
A pod of Bottlenose dolphins (Tursiops truncatus) swimming into the Sado river, Portugal, July
House sparrows (Passer domesticus) perched on building, Morocco, March
Lapwing (Vanellus vanellus) group roosting in the lagoons at Titchwell RSPB Reserve, Norfolk, UK, October. Non-ex
Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October
African elephants (Loxodonta africana) bulls crowd into an emptying waterhole to drink, Savuti, Botswana. Taken on location for BBC Planet Earth series, 2005
Black-legged Kittiwake (Rissa tridactyla) on ice. Floe edge, Arctic Bay, Nunavut, Canada, April
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Wood Ant (Formica rufa) workers on top of their nest synchronise ejection of formic acid droplets to maximise deterrent effect on potential predator, UK
Herd of Wildebeest (Connochaetes taurinus) silhouetted at dusk, Serengeti NP, Tanzania, East Africa, January
Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean
Caribbean reef shark (Carcharhinus perezi) swims over a coral reef. Grand Bahama Island, Bahamas. Tropical West Atlantic Ocean
A pod of Atlantic spotted dolphins (Stenella frontalis) riding on the bow wave of a boat. Sandy Ridge, Little Bahama Bank. Bahamas
Common / Eurasian cranes (Grus grus) small flock of juveniles taking flight over the frozen, snow covered pastureland on a foggy winter morning
Whooper swans (Cygnus cygnus) in flight at dusk, Lancashire, UK