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Two Arctic hares (Lepus arcticus) fighting, Northeast Greenland National Park, Greenland. Veolia Environnement Wildlife Photographer of the Year 2009: Mammal Behaviour category: Runner-up
European Hare (Lepus europaeus) leveret feeding on grass. Wales, UK, May
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Lioness (Panthera leo) carrying her cub, Masai-Mara Game Reserve, Kenya
Hazel Dormouse (Muscardinus avellanarius) hibernating amongst leaves. Captive. Leicestershire, UK, October
Red Fox (Vulpes vulpes) cub in late evening light, Leicestershire, England, UK, July
Wolf (Canis lupus) pack running in deep snow, captive in enclosure of the Bavarian Forest National Park, Germany, February
Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain
Female Scottish wildcat (Felis silvestris grampia) portrait, in winter coat, semi habituated living wild in native oakwood, Lochaber, Scotland, April
Eastern Grey Bamboo Lemur (Hapalemur griseus) feeding on bamboo shoot, Andasibe-Mantadia National Park, Madagascar, October
European Lynx (Lynx lynx) adult male stalking through winter birch forest, Bardu, Norway, captive
Sea Otter floating at sea surface holding pup. Alaska (Enhydra lutris) USA
Impala with Oxpeckers. Kruger National Park, South Africa
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Sea otter (Enhydra lutris) resting at the sea surface with paws in the air. Barkley Sound, Vancouver Island, Canada, August
Two Giant pandas {Ailuropoda melanoleuca} climbing tree trunk in snow storm, Sichuan, China, captive
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Red Deer stag and hind (Cervus elaphus) silhouetted at sunset, Holkham Park, Norfolk, UK, December
Three North American / Canadian Otters (Lutra canadensis) lying on each other by water. Captive, UK
Minature Shetland Pony in snow, UK
Common dolphins (Delphinus delphis) surfacing, Fisheye lens. Pico, Azores, Portugal, June 2009
European bison (Bison bonasus) behind tree, Bialowieza NP, Poland, February
Badger (Meles meles) cub amongst long grass, Dorset, England, UK, July
Red Deer (Cervus elaphus) stag seen from behind a tree. Cairngorms National Park, Scotland, February
Red Fox (Vulpes vulpes) in distance in snow habitat. Vosges, France, December
Bison (Bison bison) in snow. Yellowstone National Park, USA, February
Dromedary camel (Camelus dromedarius) train crossing Merzouga dunes, Tafilalt, Sahara, Morocco, March 2007
Aye-aye (Daubentonia madagascariensis) extracting beetle grubs from wood. Endemic to Madagascar. Photographed under controlled conditions at Durrell Wildlife Conservation Trust, Jersey, UK. captive
Young Badger (Meles meles) foraging in woodland on edge of woodland, The Black Forest, Germany, May
Amazon river dolphin (Inia geoffrensis), Rio Negro, Amazonia, Brazil
Highland cattle bull portrait {Bos taurus} Scotland UK
White domestic Alpaca {Lama pacos}, bred in the UK for its soft wool, UK
Asiatic Wild Water Buffalo (Bubalus arnee), Royal Manas National Park, Bhutan
Three banded armadillo {Tolypeutes tricinctus} in defensive ball. Captive, UK
Female Eastern grey kangaroo (Macropus giganteus) standing next to joey, Grampians National Park, Victoria, Australia, May
European rabbit (Oryctolagus cuniculus) young rabbit stands alert in grass, Norfolk, UK, June. Non-ex
Red deer stag (Cervus elaphus) calling during rut. Richmond Park, England, UK, Europe
Campbells Russian dwarf hamster (Phodopus campbelli) opal colour form
Black Jaguar or Panther (Panthera onca) snarling, captive, Peru
Portrait of a meerkat (Suricata suricatta) or suricate, looking at the camera and standing to survey its territory on the edge of Makgadikgadi Pans National Park, Botswana, April
German Shepherd / Alsatian, puppy, 4 months, sitting with tongue out
RF- European grey wolf (Canis lupus) running through snow in birch forest, Tromso, Norway. Captive, April
Grey wolf howling, USA
Fosa / Fossa (Cryptoprocta ferox) male snarling, dry deciduous forest, Kirindy Forest, Western Madagascar, IUCN vulnerable species
Two smooth coated Jack Russell Terrier puppies, tan and white, 6 weeks
Rough coated Jack Russell Terrier puppy, black and tan, portrait
Head portrait of a Giant panda (Ailuropoda Melanoleuca) Bifengxia Giant Panda Breeding and Conservation Center, Yaan, Sichuan, China