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Male Japanese giant salamander (Andria japonicus) emerging from a nest hole in order to breathe, Japan, August
Pine Grosbeak (Pinicola enucleator) male feeding on berries with traffic light behind, Oulu, Finland, December
Giraffe necked weevils (Trachelophorus giraffa) male. Endemic to Andasibe, Madagascar
Northern Cardinal (Cardinalis cardinalis) male perched on branch of Crape Myrtle (Lagerstroemia) New Braunfels, San Antonio, Hill Country, Central Texas, USA
Cheetah (Acinonyx jubatus) male yawning, Kruger national park, South Africa
African Lion (Panthera leo) head portrait of male, in the grasses of Lower Mara in the Masai Mara Game Reserve, Kenya
Killer Whale / Orca (Orcinus orca) large bull swimming in Resurrection Bay, Kenai Fjords National Park, outside Seward, Alaska, May
African Lion (Panthera leo) Samburu NP, Kenya
Leopard (Panthera pardus) male climbing on fallen tree, , Nakuru national park, Kenya
Impala (Aepyceros melampus) male, Masai-Mara Game Reserve, Kenya
Agama (Agama agama) male, Tsavo East National Park, Kenya
African Buffalo (Syncerus caffer) males fighting, Nakuru National park, Kenya
Jogi, a Jack Russell Terrier cross breed, male playing in the snow. Germany
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Highland bull (Bos taurus) face close up in freezing fog, Berwickshire Scotland, December
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Jumping spider (Euophrys frontalis) male amongst flower petals, Peak District National Park, Derbyshire, UK, June
Male Indian peafowl (Pavo cristatus) displaying, captive
Male Whinchat (Saxicola rubetra) perched on Oil seed rape (Brassica napus) in a field, Vosges, France, April
A rare Sorraia stallion cantering, Coudelaria Nacional (National Stud Farm), in Alter do Chao, District of Portalegre, Alentejo, Portugal
Jaguar (Panthera onca palustris) male wading through water at the shore of the Piquiri River. The Pantanal wetlands of Mato Grosso, Mato Grosso State, Brazil, October
African elephants (Loxodonta africana) bulls crowd into an emptying waterhole to drink, Savuti, Botswana. Taken on location for BBC Planet Earth series, 2005
Pheasant (Phasianus colchicus) male amongst Bluebell flowers, UK, April
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
Couchs Spadefoot (Scaphiopus couchii) adult at night calling with vocal sac inflated. Laredo, Webb County, South Texas, USA, April
Chestnut Paso peruvian stallion running, California, USA
Hoolock Gibbon (Hylobates hoolock) male hanging from a branch. Assam, India, February
Red Deer (Cervus elaphus) male standing in snow. The Netherlands, January
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Grey Wolf (Canis lupus) head portrait of male, sleeping in snow, Captive
Red deer (Cervus elaphus) stag amongst winter vegetation, Salburua Park, Alava, Spain, November
African lion (Panthero leo) male giving flehmen grimace, Tanzania, East Africa
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
Polar bear (Ursus maritimus) portrait of curious young male, covered in snow from rolling around / playing. On a barrier island during autumn freeze up, Bernard Spit
Harlequin duck (Histrionicus histrionicus) portrait of male stood motionless on exposed rock, in river, Iceland, May
Ptarmigan (Lagopus mutus) male in flight over snow covered Cairngorm Mountains, Scotland, February. (non-ex)
Old male African lion (Panthera leo) portrait, Sabi Sand Game Reserve, South Africa, June
Giraffe necked weevil {Trachelophorus giraffa} male on leaf of host plant (Dichaetanthera cordifolia) tropical rainforest, Mantadia NP, Madagascar
Dalmatian, head portrait, male aged 4 years
Adult male Bornean orangutan (Pongo pygmaeus) in rainforest canopy, Gunung Palung National Park, Borneo, West Kalimantan, Indonesia
HeaPair of male Boxers fawn coloured with natural ears, sitting by lake in Autumn, Illinois, USA
Portrait of male Boxer, light brindle coloured with natural ears, standing by lake shore, Illinois, USA
A Sika deer stag rests among a band of wild Misaki-uma horses (Equus ferus caballus) in the Cape Toi Reserve, Miyazaki Prefecture, Kyushu Island, Japan
A wild Misaki-uma breeding stallion (right) brings back one of his mares to his band while another breeding stallion (left) charges him away, in the Cape Toi Reserve, Miyazaki Prefecture
Male American robin (Turdus migratorius) puffed up to keep warm, and perched on Juniper branch, Kentucky, USA
Head portrait of male Mandrill (Mandrillus sphinx) Captive. Apenheul zoo; the Netherlands
Male African lion (Panthera leo) lying in grass at sunrise, Masai Mara National Reserve, Kenya, January