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Lowland gorilla (Gorilla gorilla) mother and young in forest, Loango National Park, Gabon. January
Asiatic lion (Panthera leo persica) cub, Gir Forest National Park, Gujarat, India. May
Close up of a Harp seal (Pagophilus groenlandicus) pup, Magdalen Islands, Canada. March
Coastal grey wolf (Canis lupus), British Columbia, Canada. August
RF - African lion cub (Panthera leo), age 3 months, South Luangwa National Park, Zambia (This image may be licensed either as rights managed or royalty free.)
Giant river otter (Pteronura brasiliensis) surfacing from river, Pocone, Brazil
Cleaning shrimp (Periclimenes sp) on a coral reef, close up, Triton Bay, West Papua, Indonesia, Pacific Ocean
Hawksbill turtle (Eretmochelys imbricata) cloze up on coral reef, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Green ringtail possum (Pseudochirops archeri) grooming itself on tree branch in rainforest, Atherton Tablelands, Wet Tropics World Heritage area, Queensland, Australia
Badger (Meles meles) alking through long grass, near Launceston, Cornwall, UK. July
Two Arctic foxes (Vulpes lagopus), blue-morph with winter coats, playing, Hornstrandir Nature Reserve, Iceland. March
Iceland horses in winter, western Iceland. March
Caribbean flamingo (Phoenicopterus ruber) chick, standing by its parent's legs in its nest, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Bookplate
Close up of a Caribbean flamingo (Phoenicopterus ruber) in the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Bookplate
Caribbean flamingo (Phoenicopterus ruber), group, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Young Long clawed squat lobster (Munida rugosa), portrait, Loch Linnhe, The Highlands, Scotland, UK
RF - Tompot blenny fish (Parablennius gattorugine) peering out from hole under Swanage Pier, Dorset, UK, (This image may be licensed either as rights managed or royalty free.)
Ocelot (Leopardus pardalis) hiding amongst tree roots, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Emerald damselfy (Lestes sponsa) at Ledstone pond near Leeds, UK. September. Focus stacked
Fulmar (Fulmarus glacialis) in flight against sky, Orkney, Scotland, UK, May
European badger (Meles meles) in woodland in late evening light. Scotland, UK, August
Crested tit (Lophophanes cristatus) Sierra de Grazalema Natural Park. Southern Spain. August
Himalayan ibex (Capra sibirica hemalayanus) males, resting and grazing. They live at elevations of 3800m and higher, western Himalaya mountains, Kibber Wildlife Sanctuary, India. April
Portrait of Emu (Dromaius novaehollandiae). Kangaroo Island, South Australia, Australia
Blue knee hermit crab (Dardanus guttatus) prefers shells with narrow openings such as this cone shell, Yap, Micronesia
Flame tip hermit crab (Calcinus minutus), a left-handed hermit crab that is often found in branching hard corals, Yap, Micronesia
Pygmy owl (Glaucidium passerinum) perched on branch. Helsinki, Finland. January
Great grey owl (Strix nebulosa) in snow, portrait. Kuhmo, Finland. February
Red squirrel (Sciurus vulgaris) in winter pelage, in tree. Muurame, Finland. October
Wolf (Canis lupus), young male walking through grassland. Kiekinkoski, Finland. June
RF - Red deer (Cervus elaphus) stag standing amongst Scots pine (Pinus sylvestris) tree trunks in forest. Cairngorms National Park, Scotland, UK. January
Cunner (Tautogolabrus adspersus) pair in Eelgrass (Zostera marina) bed. Newfoundland, Canada. September
Young female Grey seal (Halichoerus grypus) beneath sunburst. Farne Islands, Northumberland, UK
Dalmatian pelican (Pelecanus crispus) portrait, Lake Kerkini, Greece, February
Striped Slender Robberfly (Leptogaster cylindrica) Ledston, Yorkshire, July. Focus stacked image
Lanzas salamander (Salamandra lanzai), endemic to Cottian Alps, Monviso massif, Italy
Mute swan (Cygnus olor) cygnet swimming. Richmond Park, London, UK. May
Long-eared owl (Asio otus) looking down from a tree in autumn, Wijchen, The Netherlands, December. Second Prize Winner, Birds in their Environment Category
Arctic fox (Vulpes lagopus) cub in meadow, portrait. Hornstrandir Nature Reserve, Iceland, July
Arctic fox (Vulpes lagopus) walking in heavy snow. Hornstrandir Nature Reserve, Iceland. February
Arctic fox (Vulpes lagopus) resting in snow. Hornstrandir Nature Reserve, Iceland, February
Steller sea lions (Eumetopias jubatus) playing underwater, Inian Island, Alaska, USA. August
Red fox (Vulpes vulpes) head portrait in snow, Jura, Switzerland (Book cover image)
