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Panther chameleon (Furcifer pardalis) sitting on branch with black background, Nosy Faly, Madagascar
Marine iguana (Amblyrhynchus cristatus) swimming underwater, Fernandina Island, Galapagos
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Armadillo Lizard (Cordylus cataphractus) biting its own tail while trying to roll into a defensive ball, captive, from South Africa
Frill-neck Lizard (Chlamydosaurus kingii), displays on a termite mound. Northern Territory, Australia
Senegal chameleon (Chamaeleo senegalensis) walking over flat ground, Allahein river, The Gambia
Centralian knob-tailed gecko (Nephrurus amyae), large adult, near Alice Springs, November
Thorny devil (Moloch horridus) walking over sand dune, Yulara, Northern Territory, Australia, November
Common collared lizard (Crotaphytus collaris auriceps) male basking on rock. Arches National Park, Utah, USA. May
Thorny devil (Moloch horridus). Kalbarri National Park, Western Australia. October
Smooth knob-tailed gecko (Nephrurus levis occidentalis) portrait. Kalbarri, Kalbarri National Park, Western Australia. October
Sambava Panther Chameleon (Furcifer pardalis), Eastern Madagascer, controlled conditions
Crested gecko (Correlophus ciliatus), New Caledonia, controlled conditions
Central bearded dragon (Pogona vitticeps) male on stick in chenopod shrubland habitat, Meringur in NW Victoria, Australia
Tokay gecko (Gekko gecko) enacting a defensive display towards a perceived threat, Phuket Island, Thailand, March. Controlled conditions
Phuket horned tree lizard (Acanthosaura phuketensis) near Bang Pae Waterfall, Khao Phra Thaew NP, Phuket Island, Thailand
Portrait of an adult Three-toed Earless Skink (Hemiergis talbingoensis) Tallarook, Goulburn River in central Victoria, Australia
Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
Crocodile monitor (Varanus salvadorii) close up eye, captive, occurs in New Guinea
Deccan ground gecko (Cyrtodactylus albofasciatus) on forest floor, backlit with water droplets. Amboli, Maharashtra, India. Endemic to Western Ghats
Indonesian blue-tongued skink (Tiliqua gigas) with tongue extended, captive, from Papua New Guinea
Marine iguana (Amblyrhynchus cristatus) males in breeding colouration on rocks, with sun halo, Floreana Island, Galapagos
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
Lava lizard (Microlophus peruvianus) juvenile peering out of eye of dead sea lion. Paracas National Reserve, Peru. Highly commended in the Animals in their Environment Category of Wildlife
Slow worm (Anguis fragilis) on coastal clifftop grassland. Cornwall, England, UK. May
Tokay gecko (Gekko gecko) Shek Pik, Lantau Island, Hong Kong, China
Komodo dragons (Varanus komodoensis) on shore with saliva dripping from mouth, Rinca Island, Komodo National Park, Indonesia
Gila monster (Heloderma suspectum) Arizona, USA. Captive
Water monitor (Varanus salvator), portrait, flicking tongue, Thailand
RF - Mossy Leaf-tailed Gecko (Uroplatus sikorae) resting and camouflaged on tree trunk in rainforest understorey. Mantadia National Park, east Madagascar
Thorny dragon (Moloch horridus) in desert habitat, Australia
Texas horned lizard (Phrynosoma cornutum) portrait, Laredo Borderlands, Texas, USA. April
Nosy Be Pygmy Leaf Chameleon (Brookesia minima) female, worlds smallest reptile. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Blue-spotted dwarf iguana (Enyalioides praestabilis) on branch, Sumaco, Napo, Ecuador
Amazon wood lizard / Guichenots dwarf iguana (Enyalioides laticeps) portrait, Yasuni National Park, Orellana, Ecuador
Shieldhead gecko (Gonatodes caudiscutatus) on branch, Canande, Esmeraldas, Ecuador
Pinocchio lizard (Anolis proboscis) male on twig, Mindo, Pichincha, Ecuador, January 2013, Endangered species
Marine Iguana (Amblyrhynchus cristatus) perched on a rock. Floreana Island, Galapagos Islands, Ecuador
Namib sand gecko (Pachydactylus rangei) in sand, Swakopmund, Erongo, Namibia
Lance-nosed chameleon (Calumma gallus), Andasibe-Mantadia National Park. Madagascar
Cat gecko (Aeluroscalabotes felinus) with coiled tail, Sarawak, Malaysian Borneo
Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species
Satanic leaf-tailed gecko (Uroplatus phantasticus) on twig, Andasibe-Mantadia National Park, Alaotra-Mangoro Region, Madagascar
Parsons Chameleon (Calumma parsonii) climbing in rainforest understorey. Andasibe-Mantadia National Park, eastern Madagascar
Komodo Dragon (Varanus komodoensis) in habitat, Komodo National Park, Indonesia