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Borneo giant forest scorpion (Heterometrus longimanus) resting inside a fallen hollow log. Danum Valley, Sabah, Borneo. Photographed with natural light and UV light
Shafts of light at dusk over Wrynose pass and the Coniston hills from Ambleside, Lake District, England, UK. September 2015
Geraldton wax flower (Chamelaucium uncinatum), nectar fluorescing in UV light. Western Australia. Controlled conditions, focus stacked. Series 1/2
Kedswick in autumn, Lake District, England, UK
Kelp forest (Laminaria hyperborea) in the clear seas of Scotland, UK, September
Puffins (Fratercula arctica) Shaint Isles, Scotland, UK, June
Click beetle (Pyrophorus sp) showing orange bioluminescence, Costa Rica
Red Deer (Cervus elaphus) hind in early morning mist. Bradgate Park, Leicestershire, UK, September
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
Tropical luminous click beetle {Pyrophorus sp.} Trinidad, West Indies. Note - photographed by its own bioluminescence
Ling (Calluna vulgaris) and Bell Heather (Erica cinerea) blooming on heathland with sun rising through tree. Pig Bush, Beaulieu, New Forest National Park, Hampshire, England, UK, August
Lanternfish (Lepidophanes guentheri) - deepsea species showing bioluminescence
Limestone cave and waterfall, The Foradada, Cantonigrs, Catalonia, Spain, February 2010
Wandering Albatross (Diomedea exulans) flying over the rugged coast near Stromness, South Georgia, South Atlantic. (digitally stitched image) February
Sunrise through deciduous woodland with beech trees casting long shadows. Bolderwood, New Forest National Park, Hampshire, England, UK, October
Dawn in Bolderwood with mist and rays of sunlight. New Forest National Park, Hampshire, England, UK, September
Sheep stuck in heavy snow near Ambleside, England, UK. March 2006
Kermit pool in Hancock gorge, Karijini National Park, Pilbara, Western Australia. August 2009
Basking shark (Cetorhinus maximus) disappears back into the blue in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June
Wych Elm (Ulmus glabra) dead trees in mist at dawn, Berwickshire, Scotland, July
Black-eyed susan (Rudbeckia hirta) illuminated using different waveforms of light. Visible light (left), reflected UV light (centre), UV light as seen by bees (right). Studio environment, composite
Women walking the dog in heavy snow near Ambleside, England, UK. March 2006
Beech (fagus) forest in autumn. Vosges mountain, France, October
Manaslu range seen from Laurebinya, at sunrise, Laurebinya, Langtang - Gosaikund region, Nepal, November 2009
Sunrise over mountains, viewpoint from Tharepati, Helambu region, Nepal, November 2009
Sun shining through clouds over Lake Skadar near Murici, Lake Skadar National Park, Montenegro, May 2008
European oak (Quercus robur) with rays of sunlight, Klampenborg Dyrehaven, Denmark, September 2008
RF- Light shining through treesn in a woodland bog at dawn, Bergslagen, Sweden, June
Tufts of seaweed growing on a shallow seabed, Island of Coll, Inner Hebrides, Scotland, UK, June
Suns rays shining through dark clouds, Ballynahone Bog, County Antrim, Northern Ireland, UK, June 2011
Forest interior at dawn, The National Forest, Midlands, UK, Spring 2011
Red deer (Cervus elaphus) silhouette of hind in woodland glade at sunset, Bradgate Park, Leicestershire, England, UK, October
Red deer (Cervus elaphus) stag crossing a river, Bradgate Park, Leicestershire, England, UK, October
Fallow deer (Dama dama) stag walking at dawn, Bradgate Park, Leicestershire, England, UK, October
Brown Bear (Ursus arctos) silhouetted at dawn, Suomussalmi, Kainuu region, Finland, Europe, June 2008, Exclusive Japanese calendar rights for 2014
Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, lit by morning light, stalking. Endangered. Bandhavgarh National Park, India. Non-ex
Sandhill Cranes (Grus canadensis) in flight silhouetted against dawn light. Bosque del Apache, New Mexico, USA, November
Green aurora borealis light in night sky, Senja, Norway, February 2012
View up thorugh Licuala Fan palms (Licuala ramsayi) Licuala State Forest, Mission Beach, Queensland, Australia, December
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Termites (Isoptera) flying upwards towards a street lamp, India
The snow at Horsetail Falls begins to melt in mid-February, enabling the falls to resume once again. The clear sky and low sun angle ignite the rock wall during sundown
Sun beams passing through misty trees, Montezuma National Wildlife Refuge, New York, USA. October 2008. Highly Honored in Natures Best 2010 competition
Sundog / Parhelion (circle of light around the sun) with large ice fumerole in the foreground, McMurdo Sound, Ross Sea, Antarctica, November 2008
Sunlight creating a rainbow in the spray of the Bridalveil Falls, Yosemite National Park, California, USA, June 2008