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Strawberry poison frog (Oophaga pumilio) photographed on a white background in mobile field studio, Central Caribbean foothills, Costa Rica
Humpback whale (Megaptera novaeangliae) Umbili, a female age 8 years, breaching, Francisco Coloane Marine Park, Chiles first marine park, southeast of the Carlos III Island, Strait of Magellan, Chile
Alcedo giant tortoises (Chelonoidis vandenburghi) group resting in water to keep cool, Alcedo Volcano, Isabela Island, Galapagos
Western Santa Cruz giant tortoise (Chelonoidis porteri) portrait, Highlands, Santa Cruz Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) males in breeding colouration on rocks, with sun halo, Floreana Island, Galapagos
Green turtle (Chelonia mydas), female returning to the sea after egg laying, Puerto Pajas, Isabela Island, Galapagos
Remoras (Remora sp. ) hitchhiking on the tail of a large (13-15m) female whale sharkRemoras (Remora sp.) hitchhiking on the tail of a large (13-15m) female whale shark (Rhincodon typus) swims in open water. Wolf Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
River running through cloud forest, Manu Biosphere Reserve, Amazonia, Peru
Dobsonfly female (Corydalinae) in cloud forest, Manu Biosphere Reserve, Peru
Great white shark (Carcharodon carcharias) breaking surface with mouth open. Guadalupe Island, Mexico
Remoras (Remora sp. ) hitchhiking on the tail of a large (13-15mRemoras (Remora sp.) hitchhiking on the tail of a large (13-15m) female whale shark (Rhincodon typus) swims in open water. Wolf Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Amazonian Horned Frog (Ceratophrys cornuta) camouflaged amongst leaf litter on lowland rainforest floor, waiting to ambush passing prey. Manu Biosphere Reserve, Amazonia, Peru. November
Giant waxy monkey frog (Phyllomedusa bicolor). Lowland Amazon rainforest, Manu Biosphere Reserve, Peru
Brown / Tufted Capuchin (Cebus apella) family group in cloud forest, Manu Biosphere Reserve, Peru
Oreocallis grandiflora (Proteaceae family), Manu Cloud forest at 3500 metres altitude, Peru. November
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
White witch moth (Thysania agrippina) camouflaged against bark of tree. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
RF - Maidens veil / Bridal veil fungus (Phallus indusiatus) with indusium fully formed, on rainforest floor. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Tufted / Brown capuchin (Cebus apella), female with baby on back, sitting in tree, mid-altitude montane forest, Manu Biosphere Reserve, Peru
Dune and Red Mangrove (Rhizophora mangle), Bahia Magdalena, Baja California Peninsula, Mexico, June
Red Mangrove (Rhizophora mangle) seedling, Bahia Magdalena, Baja California Peninsula, Mexico, June
Grey whale (Eschrichtius robustus) breaching, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Wild boar (Sus scrofa) eyes glowing at dusk, introduced species in La Pampa, Argentina
Grey whale (Eschrichtius robustus) eye, Magdalena Bay, Baja California, Mexico, February. Winner in the Ocean Views category of the 2018 Natures Best Winland Smith Rice Awards
Lava lizard (Microlophus peruvianus) juvenile peering out of eye of dead sea lion. Paracas National Reserve, Peru. Highly commended in the Animals in their Environment Category of Wildlife
Whip scorpion (Heterophrynus elephas) hunting for food on a large tree root of the rainforest. Los Amigos Biological Station, Peru
Black spider monkey (Ateles chamek) calling, captive, Peru
White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru
Black spider monkey (Ateles chamek) portrait, captive, Peru
Aerial view of Amazon Rainforest, Peru, July 2015
Aerial view of Mouth of the Yavari-Mirin River entering Yavari River and Amazon Rainforest, Peru, July 2015
Aerial view of Amazon Rainforest and the Yavari River, and oxbow lake, Peru. July 2015
Golden-mantled tamarin (Saguinus tripartitus) captive
Manu Road with small waterfall, Manu National Park, Madre de Dios Peru. August 2013. Road spans from Cuzco to the Madre de Dios River and Manu
Spanish cedar (Cedrela odorata) tree, Manu National Park, Peru
Ghost glass frog (Sachatamia ilex), Costa Rica
Wild Buffons macaw x Scarlet macaw (Ara ambiguus x Ara macao) hybrid, Costa Rica
Amazon kingfisher (Chloroceryle amazona) with fish, Cuiaba, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
California condor (Gymnogyps californianus). in wild, Baja, Mexico
Jaguar (Panthera onca) male, hunting Capybara (Hydrochoerus hydrochaeris). The capybara while jumping away has cut the nose of the Jaguar with its toemail
Jaguar (Panthera onca) male, chasing a Caiman. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Tea mangrove (Pelliciera rhizophorae) seed, Pochote Estuary, Costa Rica, Vulnerable species
Magnificent hummingbird (Eugenes fulgens) female, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica