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Female Jaguar (Panthera onca palustris) with cub (estimated age 5 months), resting on a fallen tree over the Cuiaba River. Porto Jofre, Northern Pantanal, Mato Grosso State, Brazil, South America
Jaguar (Panthera onca) resting on river bank, Pantanal, Brazil
Female Jaguar (Panthera onca), captive, occurs in Southern and Central America
Black-crowned Central American squirrel monkey (Saimiri oerstedii) sitting on branch, Corcovado National Park, Osa Peninsula, Costa Rica, Vulnerable species
Aerial view of Mangrove forest, Osa Peninsula, Costa Rica, May 2017
Tea mangroves (Pelliciera rhizophorae) Pochote Estuary, Costa Rica. Vulnerable species
Tea mangrove (Pelliciera rhizophorae) flower, Pochote Estuary, Costa Rica, Vulnerable species
Cloud forest of the Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica. Small repro only
Puma (Puma concolor) resting, Corcovado National Park, Costa Rica, May
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Bairds tapir (Tapirus bairdii) walking along beach in Corcovado National Park, Costa Rica. Endangered
Ocelot (Leopardus pardalis) walking, camera trap image, Nicoya Peninsula, Costa Rica
Ocelot (Leopardus pardalis) portrait. camera trap image, Nicoya Peninsula, Costa Rica
Leaf mantis (Choeradodis sp. ) Tortuguero National Park, Costa RicaLeaf mantis (Choeradodis sp.) Tortuguero National Park, Costa Rica
Jaguar (Panthera onca) camera trap image, Tortuguero National Park, Costa Rica
Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica
Eyespots on wing of Giant owl butterfly, (Caligo memnon) Tortuguero National Park, Costa Rica, February 2015
Neotropical river otter (Lontra longicaudis) in water, Nicoya Peninsula, Costa Rica, March 2015
Long beaked common dolphin (Delphinus capensis) pod porpoising Baja California, Mexico
Californian Sealion (Zalophus californianus) in kelp (Macrocystis pyrifera) forest, San Benito del Este Island, Baja California Peninsula Pacific Islands Biosphere Reserve, Baja California, Mexico
Ants (Formicidae) attacking newly emerged cicada (Cicadacae), Yasuni National Park, Ecuador. Highly commended in the Invertebrates Category of the Wildlife Photographer of the Year Awards (WPOY)
Water lilies in the Cenote Nicte Ha, Quintana Roo, Yucatan Peninsula, Mexico. May. Photographed for The Freshwater Project
Water hyacinth (Eichhornia sp. ) and red Ludwigia (Ludwigia sp. ) in a clear stream, Pantanal, BrazilWater hyacinth (Eichhornia sp.) and red Ludwigia (Ludwigia sp.) in a clear stream, Pantanal, Brazil. Photographed for the Freshwater Project
Piraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp. ) Rio Olhio d AguaPiraputanga (Brycon hilarii) and small Tetra fish (Hyphessobrycon sp.) Rio Olhio d Agua, tributary of Rio da Prata, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Brown pelican (Pelecanus occidentalis) feeding chicks at nest, Puerto Ayora / Academy Bay, Santa Cruz Island, Galapagos
Andean flamingo (Phoenicoparrus andinus) feeding on shore Laguna Hedionda, Altiplano, Bolivia
Jamess flamingo (Phoenicoparrus jamesi) flock on the shore of Laguna Colorada / Reserva Eduardo Avaroa, Altiplano, Bolivia
Guanaco (Lama guanicoe) with lake and mountains, Torres del Paine National Park, Patagonia. Chile
Wire-crested thorntail hummingbird (Discosura popelairii) Sumaco, Napo, Ecuador
Laguna Miluni a reservoir fed by glacial melt water from the Andean peak of Huayna Potosi. As climate change causes the glaciers to melt, the water supply for La Paz
Colourful lakes below the peak of Chacaltaya, lake discoloured by mine effluent. Andes, Bolivia. October 2015
White tent making bat (Ectophylla alba) roosting in tree, La Selva Field Station, Costa Rica
Coscoroba swan, (Coscoroba coscoroba) family with chicks, La Pampa, Argentina
Green-crowned brillian-t hummingbird (Heliodoxa jacula) visiting Torch ginger flower (Etlingera elatior), Poas Volcano National Park Costa Rica
Peruvian mantis (Oxyopsis peruviana) silhouette with sun behind, captive, occurs in South America
Scarlet Macaw (Ara macao) preening tail feather, Pantanal, Brazil
Amazon river dolphin (Inia geoffrensis) Rio Negro, Amazonas, Brazil
Galapagos sea lion (Zalophus wollebaeki) hunting tuna. A group of the sea lion bulls have learnt to herd Pelagic yellowfin tuna into a small cove, trapping them
Galapagos sea lions (Zalophus wollebaeki) hunting tuna. A group of the sea lion bulls have learnt to herd Pelagic yellowfin tuna into a small cove, trapping them
White-bellied parrots (Pionites leucogaster xanthomeri) in rainforest, Tambopata National Reserve, Peru
Leafcutter ant (Atta cephalotes) carrying pieces of leaves, Costa Rica
Vicuna (Vicugna vicugna) mother with young, Laguna Hedionda, Altiplano, Bolivia
Salar de Uyuni salt flat at sunset, Altiplano, Bolivia
Salar de Uyuni salt flat just after sunset, Altiplano, Bolivia
Spotfin Burrfish (Chilomycterus reticulatus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
RF - Reticulated Glass Frog (Hyalinobatrachium valerioi) backlit showing highly translucent body. Osa Peninsula, Costa Rica