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King penguins (Aptenodytes patagonicus) crossing water to reach breeding site, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Chilean flamingos (Phoenicopterus chilensis) in flight over mountain peaks with glacier in the distance, Torres Del Paine National Park, Chile
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Row of Larch trees on flowering heather moorland, Lammermuir Hills, Berwickshire, Scotland, August
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Mixed flock of Snow geese (Chen caerulescens atlanticus / Chen caerulescens) and Sandhill cranes (Grus canadensis) at wintering area at dawn, with full moon, Bosque del Apache, New Mexico, USA
Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on trunk of Sweetgum tree (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Wildebeest (Connchaetes taurinus) running down bank of the Mara River during migration, Masai Mara, Kenya
Whale Shark (Rhincodon typus) accompanied by Pilot Fish (Naucrates ductor) and Remora. The biggest fish in the world has only started to visit the Azores in the past three years due to water
A large herd of Tien Shan Argali (Ovis ammon karelini) making tracks through snow. Naryn National Park, Kyrgyzstan, Central Asia, November
Aylesbury ducks following in a line on village green, Weedon, Buckinghamshire, UK, October
King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia
Large group of Sunflower sea stars (Asterias / Pycnopodia helianthoides) covering rock, pacific coast, Canada, August
Knot (Calidris canutus) and grey plover (Pluvialis squatarola) roost at high tide with numerous birds packed tightly together, Snettisham, Norfolk, October
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
Flamingos on Laguna / Lake Colorado at dawn, Eduardo Avaroa Andean Fauna National Reserve, Bolivia, December 2009
Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
Common / Eurasian crane (Grus grus) flock in flight with snow topped Pyrenees mountains in the background, during southward migration period, Haute Pyrenees, Gascony, France. December 2009
Kittiwake (Rissa tridactyla) flock feeding in from of the Monaco Glacier, Leifdefjord, Svalbard, Norway 2010
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
American Buffalo / Bison (Bison bison) grazing in open plains. Yellowstone National Park, Wyoming, October
Domestic cows (Bos taurus) and free roaming horses grazing in the Pyrenees-Atlantiques, Pyrenees, France, June
A flock of Greylag Goose (Anser anser) flying in formation above misty fields. The Netherlands, March
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Little Auks (Alle alle) flying low above surface in front of iceberg. Spitsbergen, Svalbard, Norway, July 2010. Highly commended in the Birds category in the GDT competition 2011
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
King penguin (Aptenodytes patagonicus) chicks in mass breeding colony, St Andrews Bay, South Georgia. Taken on location for BBC Frozen Planet series, 2008
King penguins (Aptenodytes patagonicus) on beach, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Bigeye Scad (Selarcrumenophthalmus). Misool, Raja Ampat, West Papua, Indonesia
Barrel Sponge (Xestospongia testudinaria) on coral reef with soft corals (Scleronephthya sp), gorgonian and Lyretail anthias (Pseudanthias squamipinnis). Komodo National Park, Indonesia
Great white shark (Carcharodon carcharias) portrait, Guadalupe Island, Mexico, Pacific Ocean
Flock of Kelp / Southern black backed Gulls (Larus dominicanus) flying over iceberg. Graham Passage, Antarctic Peninsula, Antarctica. February. (digitally altered image)
Flock of Sanderling (Calidris alba) flying over a rough sea, Berwickshire Coast, Scotland, UK, March
King Penguin (Aptenodytes patagonicus) adult standing amongst colony of chicks, Gold Harbor, South Georgia, South Atlantic Islands
Galapagos garden eels (Heteroconger / Taenioconger klausewitzi) on seabed, Galapagos Islands
A bull California sealion (Zalophus californianus) bursting through a school of baitfish in the shallows. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean
Arizona mountain kingsnakes (Lampropeltis pyromelana) Found in Chiricahua mountains, SE Arirona. Controlled conditions
Wildebeest (Connochaetes taurinus) migration, herd crossing the Mara river, Masai-Mara Game Reserve, Kenya
Herd of Southern giraffes (Giraffa camelopardalis giraffa) and Burchells zebras (Equus quagga) aerial view, Okavango delta, Bostwana
Shoal of Longfin bannerfish (Heniochus acuminatus) Maldives, Indian Ocean
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Monarch butterfly (Danaus plexippus) hibernating, Mariposa Monarca Special Biosphere Reserve, central Mexico, January
American Flamingo (Phoenicopterus ruber), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August
Emperor angelfish (Pomacanthus imperator) with soft corals and sweepers. Egypt, Red Sea
