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King penguins (Aptenodytes patagonicus) crossing water to reach breeding site, South Georgia. Taken on location for BBC Frozen Planet series, 2008
The Dark Hedges, Stranocum, Co. Antrim, Northern Ireland, September 2010
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October
Track through fir forest in autumn, Vosges mountain, France, October
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
RF- Adelie Penguin (Pygoscelis adeliae) on iceberg. Yalour Islands, Antarctic Peninsula, Antarctica. February
Lime Avenue a famous row of Lime trees (Tilia Sp) Hampstead Heath, London, England, UK. October 2014
RF - Giant panda (Ailuropoda melanoleuca) in snow, captive
RF - Mute swan (Cygnus olor) taking off from a pond Valkenhorst Nature Reserve, Valkenswaard, the Netherlands June
Silky shark (Carcharhinus falciformis), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba, January. Vulnerable
People trekking with Dromedary camels (Camelus dromedarius) through chalk rock formations created by sandstorms, White desert, Sahara, Egypt, February 2009. No release available
Track / path running through snow covered Larch (Larix sp) forest in winter, Vosges, France, December
RF - American bisons (Bison bison) in Grand Teton National Park. winter. Wyoming, USA
Elephant (Loxodonta africana) female and calf under tree in the rain, Masai-Mara game reserve, Kenya
The road to Okeford Fitzpaine; dawn on Midsummers Day, in the Blackmore Vale, from Okeford Hill, Dorset, England, UK. June 2019
Three Sockeye salmon/ Red salmon (Oncorhynchus nerka) swimming upstream as they migrate back to the river of their birth to spawn
Two people walking in vast landscape with dry grass Barguzin Valley, Buryatial, Siberia, Russia. April 2016
Aerial view of two Gemsbok (Oryx gazella) in sand dunes, Namib Desert, Namibia
Great crested grebe (Podiceps cristatus) close-up of an adult with two young chicks. The Netherlands. June 2014
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
African elephant (Loxodonta africana) herd walking in procession, Kaokoveld Desert, Namibia
American bison (Bison bison) walking through snow using tracks made by other animals, Yellowstone National Park, Wyoming, USA, February
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Loggerhead turtle (Caretta caretta) hatchling running to the sea, lycian coast, Mediterranean Sea, Turkey
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Avenue of Oak trees (Quercus) Eickelberg, Warnowtal, Germany, September
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
RF- Rear view of male Bengal tiger (Panthera tigris tigris) walking on track. Bandhavgarh National Park, India. Endangered species
Leatherback turtle (Dermochelys coriacea) female, returning to sea after laying eggs in nest on beach, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Two Leatherback turtle (Dermochelys coriacea) hatchlings moving across beach towards sea after leaving nest, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean Sea
Leatherback turtle (Dermochelys coriacea) hatchling, swimming vigorously across a rainwater pool toward the sea just after leaving nest, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean
Green sea turtle (Chelonia mydas), hatchling, making its way across the beach to the ocean, Yap, Micronesia, Pacific Ocean
Female Olive ridley turtle (Lepidochelys olivacea) coming ashore at dusk to lay eggs, Oaxaca, Mexico, Pacific Ocean
Female Olive ridley turtles (Lepidochelys olivacea) coming ashore at night in large numbers to lay eggs during arribada mass nesting, Oaxaca, Mexico
Atlantic mobula ray (Mobula hypostoma) group swimming in Caribbean Sea. Isla Mujeres, Quintana Roo, Mexico
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
RF- Arabian Oryx (Oryx leucoryx) Dubai Desert Conservation Reserve, Dubai, UAE
Polar bear (Ursus maritimus) walking along coast of Wrangel Island, Far Eastern Russia, October
Polar bear (Ursus maritimus) walking in snow, Svalbard, Norway, April
Double rainbow over a road in Western Australia, December 2013
RF -Three Bison (Bison bison) walking through snow with herd feeding in background. Yellowstone National Park, Wyoming, USA. January
RF - Coyote (Canis latrans) in snow, Yellowstone. February