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Portuguese man o war (Physalia physalis) Tenerife, Canary Islands. January
Termite hill and acacia tree, Shaba National Reserve, North Kenya
Group of Bigfin reef squids (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Bigfin reef squid (Sepioteuthis lessoniana) swimming at night, Indonesia, Sea of Flores
Day octopus (Octopus cyanea) female, in mid-water, Hawaii
Rosemary beetle (Chrysolina americana) feeding on Lavender (Lavandula angustifolia) in garden Cheshire, UK, June
Queen conch (Aliger gigas) in a seagrass (Thalassia testudinum) meadow, Bahamas
Red sea fan (Paramuricea clavata) with gorgonian corals and Light-bulb sea squirt (Clavelina lepadiformis) Punta Carena, Capri Island, Sorrentine Peninsula, Italy, Tyrrhenian Sea, Mediterranean
Crinoid or feather star (Antedon mediterranea) on Violescent sea whip or Red sea fan (Paramuricea clavata) Punta Campanella Marine Protected Area, Amalfi Coast, Italy, Tyrrhenian Sea, Mediterranean
Slate pencil sea urchins (Heterocentrotus mammillatus) on coral reef with Pyramid butterflyfish (Hemitaurichthys polylepis), Hawaii
Sea anemone (Calliactis parasitica) usually associated with hermit crabs, here on a Mediterranean bonnet shell (Semicassis granulata), Ponza island, Italy, Tyrrhenian Sea, Mediterranean
Bordered patch butterfly (Chlosyne lacinia) on Indian blanket (Gaillardia pulchella). Hill Country, Texas, USA
Four-spotted chaser dragonfly (Libellula quadrimaculata), two resting on reeds. Cornwall, England, UK. May
Apollo butterfly (Parnassius apollo) female laying eggs on branch. Parainen, Finland. July
Wolf spider (Pardosa amentata) close up, Leeds, Yorkshire, England, UK, April. Focus stacked image
RF - Small tortoiseshell butterfly (Aglais urticae) nectaring on Verbena (Verbena sp). Cornwall, England, UK. September. (This image may be licensed either as rights managed or royalty free.)
Atlantic blue crab (Callinectes sapidus) hunting in Turtlegrass (Thalassia testudinum) seagrass bed, view towards sky. Florida Keys, Florida, USA
Orange and white soft corals, Dead mans fingers (Alcyonium digitatum) thriving beneath kelp (Laminaria hyperborea). St Abbs, Eyemouth, Berwickshire, Scotland, United Kingdom. North Sea
Cuttlefish (Sepia officinalis) male hovers over sea oak (Halidrys siliquosa). Babbacombe, Torquay, Devon, England, United Kingdom. English Channel. North East Atlantic Ocean
RF - Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and Sea plumes (Pseudopterogorgia sp)
Caribbean reef squid (Sepioteuthis sepioidea) eating fish at night, portrait. The Bahamas
White and yellow nudibranch (Polycera quadrilineata) feeding on a sea mat (Membranipora membranacea). Shetland Islands, Scotland, British Isles. United Kingdom. North East Atlantic Ocean
Lord Howe Island stick insect (Dryococelus australis), Bristol Zoo Gardens, Bristol, UK
Apple hawk-moth (Langia zenzeroides formosana) Nzaus Mamei Jianshih Village, Hsssinchu County, Taiwan, Controlled conditions
Crab spider (Thomisus onustus) on white flowerhead, Sierra de Grazalema Natural Park, southern Spain, May
Spoonwing / Thread-winged antlion (Nemoptera bipennis) on flower, Sierra de Grazalema Natural Park, southern Spain. June
Cockchafer beetle (Melolontha melolontha) on lichen encrusted twig. France. May
Firefly (Lamprohiza splendidula) light trails in woodland at night, multiple exposure. The Netherlands. June
Portrait of a Emerald damselfly (Lestes sponsa) Yorkshire, August. Focus stacked image
Stag beetle (Lucanus cervus), adult male on oak tree, Italy
Black-veined white (Aporia crataegi) butterfly pair, in flight and nectaring on Thistle. Jyvaskyla, Central Finland. June
A reef scene with an orange elephant ear sponge (Ianthella basta) and red sea whip (Ellisella sp) with schooling silversides fish above. Pelee Island, Misool, Raja Ampat, West Papua, Indonesia
Split level photo of mangrove scenery, with hard corals ( including Goniopora sp.; Heliopora sp; Porites sp.) growing below Red mangrove tree: (Rhizophora mangle)
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
RF - Open brain coral (Trachyphyllia geoffroyi) lives in symbiosis with zooxanthellae which are responsible for nutrition and the vibrant colours. Lembeh Strait, North Sulaweisi, Indonesia
RF - Male Comet Moth (Argema mittrei). Mantadia National Park, eastern Madagascar. (This image may be licensed either as rights managed or royalty free.)
Brimstone butterfly (Gonepteryx rhamni) resting among foliage, Meeth Quarry, Devon, UK. June
Marbled White butterfly (Melanargia galathea) resting on flower head, Dunsdon Nature Reserve, Devon, UK. July
Male common blue butterfly (Polyommatus icarus) basking wings open on grass, Vealand Farm, Devon, UK. June
Glasswing butterfly (Greta oto), Mariposario del Drago, Icod de Los Vinos, Tenerife, August
Painted lady butterfly (Vanessa cardui) feeding on knapweed Sutcliffe Park Nature Reserve, Eltham, London, England, UK. June
Inser Weiher in autumn, a pond with red coloured water, caused by dissolved tannins. Berner Seeland, Berne, Switzerland, November. Photographed for The Freshwater Project
Army ant (Eciton burchellii) carrying prey, Copalinga Reserve, Ecuador
Giant Pacific octopus (Enteroctopus dofleini) showing arms and suckers, Vernon Rock, Queen Charlotte Strait, British Columbia, Canada. September
A Giant rock scallop (Crassadoma gigantea) encrusted with and surrounded by several other invertebrates animals, including White anemones (Metridium sp.), White mushroom ascidians (Distaplia sp.)
Gooseneck barnacles (Pollicipes Polymerus) amongst kelp, Nakwakto Rapids, Slingsby Channel, British Columbia, Canada. September
Purple and Ochre sea stars (Pisaster ochraceus) preying on Pacific blue mussels (Mytilus trossulus). Also seen are Green Surf Anemones (Anthopleura xanthogrammica)
Stubby squid (Rossia pacifica) Nigei Island, Queen Charlotte Strait, British Columbia, Canada