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Slender sapsucking slug (Thuridilla gracilis) on surface of Royal seasquirt (Polycarpa aurata) Aljui Bay, Waigeo Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Pink anemonefish (Amphiprion perideraion) in Purple magnificent sea anemone (Heteractis magnifica) Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Harlequin shrimp (Hymenocera elegans) on blue sea star. This species of shrimps feeds on starfish, turning them upsidedown and keeping them as living larder. Bitung, North Sulawesi, Indonesia
Chocolate-dip chromis (Chromis dimidiata) male over out of focus hard coral (Porites nodifera). Sha ab Claudia, Fury Shoal, Egypt. Red Sea
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon, on coral reef under stormy seas. St Johns Reef. Egypt. Red Sea
Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea
Aggregation of tiny isopods (Santia sp. ) living on the surface of a Blue sponge (Haliclona sp)Aggregation of tiny isopods (Santia sp.) living on the surface of a Blue sponge (Haliclona sp.), there are two colour varieties here, possibly different species
RF - Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Studio image of multiple Nudibranch species. Gulen, Norway. North East Atlantic Ocean. Digital composite
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
Bigfin reef squid (Sepioteuthis lessoniana) capturing a pelagic shrimp with long red antennae, in midwater, at night. Anilao, Batangas, Luzon, Philippines
Coleman shrimp (Periclimenes colemani) close up, on a fire urchin (Asthenosoma varium). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Purple short tentacle mushroom coral (Fungia sp. ) close up of polyps. Lembeh StraitPurple short tentacle mushroom coral (Fungia sp.) close up of polyps. Lembeh Strait, North Sulawesi, Indonesia
Pink anemonefish (Amphiprion perideraion) in a purple Magnificent sea anemone (Heteractis magnifica). Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Large white butterfly (Pieris brassicae) feeding on lavender flowers in garden, England, UK, August
Starfish close up detail, Blairgowrie, Mornington Penisular, Victoria, Australia. Port Philip Bay, Bass Strait
Garden snail (Helix aspersa) baby on Marigold flower, UK
Close-up of the polyps out of a coral tree (Tubastraea coccinea) at night, Baja California peninsula, Mexico. Sea of Cortez
Orange cup coral (Tubastrea coccinea) Malpelo Island National Park, UNESCO World Heritage Site, Colombia, East Pacific Ocean
Red Admiral Butterfly (Vanessa atalanta) on Michaelmas daisy flowers. Dorset, UK, October 2012
Brimstone moth (Opisthograptis luteolata) Banbridge, County Down, Northern Ireland, June
Chinese moon moth (Actia sinensis subaurea) occurs in Northern China
Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
RF - Marbled White butterfly (Melanargia galathea) resting on common spotted orchid, Dunsdon Nature Reserve, Devon, UK. July
Emperor angelfish (Pomacanthus imperator) swimming past soft corals (Dendronephthya sp]), Red Sea, Egypt. January
Flatworm (Maiazoon orsaki) Raja Ampat, Irian Jaya, West Papua, Indonesia, Pacific Ocean
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
RF- Boulder star coral (Montastrea annularis) showing fluorescent green coloration, photographed under blue light at night, to excite fluorescent pigments in the coral, East End, Grand Cayman
Yellow springtails (Symphypleona) on Forget-me-not flower (Myosotis) Zillertal Alps, Austria, July
Common blue butterfly (Polyommatus icarus) female, Southwest London, England, UK. June
Morpho butterfly (Morpho peleides) photographed on white background. Costa Rica, March
Blue sea star (Linckia laevigata) on coral reef, Fiji, South Pacific
RF - Long-spined sea urchin (Astropyga radiata). Lembeh Strait, North Sulawesi, Indonesia
Gorgonian coral (Acanthogorgia sp. ) Similan Islands, Andaman Sea, ThailandGorgonian coral (Acanthogorgia sp.) Similan Islands, Andaman Sea, Thailand
Scarlet lily beetle (Lilioceris lilii) Nottinghamshire, England, UK, May. Introduced species
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April
Brown-lipped snail (Cepaea nemoralis) on fungus, reproduced in a bokeh affect, Sheffield, England, UK. November. Commended in the Natures Studio Category of GDT competition 2014
Female Tassled wobbegong shark (Eucrossorhinus dasypogon) rests alongside stand of Barrel sponges (Xestospongia testudinaria) Blue Magic, Raja Ampat, West Papua, Indonesia
Reef octopus (Octopus cyanea) emerges from its den in a hole in a large hard coral (Porites sp)Reef octopus (Octopus cyanea) emerges from its den in a hole in a large hard coral (Porites sp.) at sunset. Sha ab Mahmood, Sinai, Egypt. Red Sea
RF- Secretary blenny (Acanthemblemaria maria) peering from hole in massive Starlet coral (Siderastrea siderea). East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Leaf cutter ants (Atta sp) female worker ants carry pieces of fern leaves to nest, reflected in water, Costa Rica, December
Pelagic mollusc (Oxygyrus keraudreni), captive deep sea species from Atlantic Ocean off Cape Verde
Marine planktonic Copepod (Sapphirina sp.) brightly coloured copepods. Deep sea species from Atlantic Ocean off Cape Verde. Captive