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Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Leaf cutter ants (Atta sp) carrying colourful plant matter, reflected in water, Laguna del Lagarto, Santa Rita, Costa Rica
Common blue butterflies (Polyommatus icarus) basking in the morning light, Vealand Farm, Devon, UK. July
Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Acorn weevil (Curculio glandium) Portrait on an Oak bud, Hertfordshire, England, UK, May - Focus Stacked - Captive
Malaysian Orchid Mantis (Hymenopus coronatus) pink colour morph, camouflaged on an orchid. Captive, from Malaysia
Trap-jaw ant (Odontomachus chelifer) portrait, Wayqecha, Peru
Mating pair of chalkhill blue butterflies (Lysandra coridon) with wings closed resting on Devils-bit scabious (Succisa pratensis), Hatch Hill, Somerset, UK. August
Malaysian Orchid Mantis (Hymenopus coronatus) white colour morph camouflaged on an orchid. Captive. Originating from Malaysia
RF - Composite image of tropical nudibranchs on a black background showing variety and abundance of nudibranch species, Indo-Pacific
Deep sea jellyfish (Periphylla periphylla) juvenile, Trondheimsfjord, Norway, Atlantic ocean
Borneo giant forest scorpion (Heterometrus longimanus) resting inside a fallen hollow log. Danum Valley, Sabah, Borneo. Photographed with natural light and UV light
Small pearl-bordered fritillary (Boloria selene) butterfly on oxeye daisy (Leucanthemum vulgare), Marsland mouth, North Devon, UK. May 2017
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Glanville fritillary butterfly (Melitaea cinxia) roosting on larval foodplant Ribwort plantain (Plantago lanceolata), UK - Captive - Focus Stacked Image
Water fleas (Daphnia sp. ) and a green algae (Volvox aureus) in water from a garden pondWater fleas (Daphnia sp.) and a green algae (Volvox aureus) in water from a garden pond. Derbyshire, UK. September. Digital composite
Fen raft spider / Great raft spider (Dolomedes plantarius) sub-adult. Norfolk Broads, UK, September. Vulnerable species
Goldenrod crab spider (Misumena vatia) yellow female on Honesty flower, Bristol, UK, April
Electric fileclam / Disco clam (Ctenoides ales). West Papua, Indonesia. Indo-West Pacific
RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed. These octopus regularly carry pieces of shell so they can use them as protective
Nudibranch (Phyllidia marindica), Lembeh Strait, North Sulawesi, Indonesia. December
European swallowtail butterfly (Papilio machaon gorganus) on flower, Mercantour National Park, Provence, France, June
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise
RF - Devils flower mantis (Idolomantis diabolica) male, captive, occurs in Africa
Elephant hawk moth (Deilephila elpenor) portrait, Brackagh Moss National Nature Reserve, Portadown, County Armagh, Northern Ireland. April
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Edwards Atlas Moth (Attacus edwardsii) in defensive posture, Bhutan, June
Tropical shield mantis (Choeradodis sp. ) (Possibly C. rhombicollis) Canande, EsmeraldasTropical shield mantis (Choeradodis sp.) (Possibly C. rhombicollis) Canande, Esmeraldas, Ecuador
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Green tree ants (Oecophylla smaragdina) defending their leaf nest, Daintree River, Wet Tropics World Heritage area, Queensland, Australia
Portuguese man o war (Physalia physalis) Tenerife, Canary Islands
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Cairns birdwing butterfly (Ornithoptera euphorion) male resting on leaf. Kuranda Butterfly Sanctuary, Queensland, Australia. Captive
European honey bee (Apis mellifera) feeding on ice plant flowers (Sedum spectabile), Monmouthshire, Wales, UK. September
Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
Squid (Abraliopsis atlantica) deep sea species from Atlantic Ocean off Cape Verde. Captive
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
Calleta silkmoth (Eupackardia calleta), Texas USA Controlled conditions
RF - White-legged damselfly (Platycnemis pennipes) peering over Oxeye daisy (Leucanthemum vulgare) flower. The Netherlands. August
Small Tortoiseshell Butterfly (Aglais urticae) feeding on scabious flowers in hay meadow, Wensum Valley, Norfolk, England, UK, July