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Sir David Attenborough with mountain gorillas, on location for BBC series Life on Earth, Rwanda, Africa 1978
Pair of European grey wolves (Canis lupus) interacting, Tromso, Norway, captive, April
Sir David Attenborough with mountain gorillas on location for BBC series Life on Earth, Rwanda, Africa 1978
African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012
European grey wolf (Canis lupus) interacting with a European brown bear (Ursus arctos) Kuhmo, Finland, July 2009. WWE INDOOR EXHIBITION
Two white horses of the Camargue, head to head at water, Camargue, Southern France
Suffolk Punch heavy horse in field resting head anothers back, UK, September
Dolphin trainer interacting with Bottlenose Dolphin (Tursiops truncatus), Dolphin Reef, Eilat, Israel - Red Sea. Model released
Great crested grebe (Podiceps cristatus) pair performing the weed dance, a spectacular part of their elaborate courtship ritual, Derbyshire, UK, March
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Wild boar (Sus scrofa) mother watching over tiny piglets, the Netherlands, April
Goldfinches (Carduelis carduelis) squabbling over teasel seeds in winter. Hope Farm RSPB reserve, Cambridgeshire, England, UK, February
Domestic sheep, lambs playing in a field, Norfolk, UK, March
European rabbit (Oryctolagus cuniculus) two young rabbits, or kittens, touch noses, Norfolk, UK, June. Non-ex
Baby African elephants (Loxodonta africana) of various ages playing together, Addo national park, Eastern Cape, South Africa, January
Domestic black pig {Sus scrofa domestica} smelling another pigs ear, Spain
Two white horses of the Camargue, interacting, greeting head to head in marshes, Camargue, Southern France
Northern stone crab (Lithodes maja) pair, Trondheimsfjorden, Norway, February 2009
Konik horse (Equus caballus) herd grazing with two stallions interacting, Wicken Fen, Cambridgeshire, UK, June 2011
Konik horse (Equus caballus) pair interacting, wild herd in rewilding project, Wicken Fen, Cambridgeshire, UK, June 2011
Baby Grey Squirrel (Sciurus carolinensis) kissing a tabby kitten
Pheasant {Phasianus colchicus} males fighting over territory, Norfolk, UK
Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
Dalmation dogs playing on beach, Norfolk, UK, August
A pair of Australian Sea Lions (Neophoca cinerea) socialising in the shallows. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
Free diver swimming amongst Californian sealions (Zalophus californianus) Anacapa Island, California, Pacific Ocean Model released
Grey seal {Halichoerus grypus} two adolescents interact among the breaking waves, Lincolnshire, UK
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Wild Chincoteague (Equus caballus) two breeding stallions greeting one another, Chincoteague National Wildlife Refuge, Chincoteague Island, Virginia, USA, June
Black-tailed Prairie Dogs (Cynomys ludovicianus), two pups (right) seem to be kissing their parent. The behaviour is a gesture of recognition
Domestic sheep, two lambs play head-butting in a field, Norfolk, UK, March
Grey seal (Halichoerus grypus) biting divers glove, Lundy Island, Bristol Channel, England, UK, May
Moose (Alces alces) female and male covered in light dusting of snow in forest, Grand Teton NP, Wyoming, USA, October
Gentoo penguin (Pygoscelis papua) at nest with chick, Macquarie Island, Southern Atlantic, Australian Antarctica, November
Stellers sea eagle (Haliaeetus pelagicus) three eagles beside Lake Kuril, Kamchatka, Far East Russia, January
Two White stork (Ciconia ciconia) chicks, Prypiat area, Turov, Belarus, June 2009
Two Muskox (Ovibos moschatus) head to head, Dovrefjell National Park, Norway, February 2009. WWE BOOK
Two Muskox (Ovibos moschatus) face to face, Dovrefjell National Park, Norway, February 2009
Two Muskox (Ovibos moschatus) face to face, Dovrefjell National Park, Norway, February 2009 WWE BOOK COVER. WWE OUTDOOR EXHIBITION. WWE INDOOR EXHIBITION. *PRESS IMAGE
Two Konik horse foals mutual grooming, Oostvaardersplassen, Netherlands, June 2009 WWE BOOK
Two Polar bears (Ursus maritimus) interacting, Svalbard, Norway, September 2009
Gannets (Morus bassanus) Saltee Islands, County Wexford, Ireland, June 2009
Urban Red fox (Vulpes vulpes) juvenile and adult, London, May
Konik horse and young foal interacting, Oostvaardersplassen, Netherland, June 2009
Red squirrel (Sciurus vulgaris) approaching another as it eats a nut, Cairngorms National Park, Scotland, March 2012
Konik horse (Equus caballus) mutual grooming, wild herd in rewilding project, Wicken Fen, Cambridgeshire, UK, June 2011