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RF - St Michaels Mount and old causeway at sunrise, Marazion, Cornwall, UK. October 2015
Bulb tentacle sea anemone (Entacmaea quadricolor) close up detail, Maldives, Indian Ocean
Pink abalone (Haliotis corrugata) shell detail, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, January
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
RF- Aerial view of the Okavango delta, Botswana, May 2010
Nosy Be Pygmy Leaf Chameleon (Brookesia minima) female, worlds smallest reptile. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Fork-leaved sundew (Drosera bipinata) leaves unfurling. Cultivated species, occurs in Australia and New Zealand
Wild Garlic / ramsons (Allium ursinum) leaves emerging in early spring. Peak District National Park, Derbyshire, UK. March
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Caribbean flamingo (Phoenicopterus ruber) about to take off to roosting site, Ria Celestun Biosphere Reserve, Yucatan Peninsula Mexico, January
Underwater scene where light and water interplay on a shallow sand bank, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea, September 2005
Five newly hatched chicks in a row
Birch tree (Betula sp) in autumn, blurred, Cairngorms National Park, Scotland, November. Winner of the Wild Woods Category of BWPA competitions 2014
Large group of Sunflower sea stars (Asterias / Pycnopodia helianthoides) covering rock, pacific coast, Canada, August
RF - Reticulated Glass Frog (Hyalinobatrachium valerioi) backlit showing highly translucent body. Osa Peninsula, Costa Rica
Blue jellyfish (Cyanea lamarckii), feeding on small plankton, Lundy Island Marine Conservation Zone, Devon, England, UK, May
School of Bluespine unicornfish (Naso unicornis) gathered during the summer breeding season. Yolanda Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
American Purple Gallinule (Porphyrio martinica) displaying brightly coloured wings. Everglades National Park, Florida, USA, March
Pyrenean chamois {Rupicapra rupicapra pyrenaica} Pyrenees, France
Water lily (Nymphaea rudgeana) leaves viewed from below in a black water creek. Suriname, October
Detail in frozen waterfall. Selijalandfoss waterfall, Southern Iceland, Europe, March 2011
Aerial image showing sandbanks in the Bahamas archipelago, Caribbean, February 2012
Leaves floating on water with colourful reflections, Plitvice Lakes National Park, Lika, Croatia, Europe, October 2011
Looking down at Chubb (Leuciscus cephalus) at surface with reflection of bridge, Rhine River, Switzerland, 1993. No release available
Aerial view of the rivers and saltmarshes of the Bahia / Bay de Cadiz Natural Park, Andalucia, Spain, March 2008
Looking up at Beech wood canopy (Fagus sylvatica) in autumn, Peak District National Park, Derbyshire, UK, November
Aerial view of Lake Natron, Rift Valley, Tanzania, Africa, August 2009
Reflections in autumn leaves in lake, Acadia National Park, Maine, USA, October
Espanola mockingbird (Mimus macdonaldi) on beach, Galapagos
Portrait of a rare five-day old Posavina filly hidden by her mothers tail, Lonja, Croatia, August
Reflection in fen on surface of water, Klein Schietveld, Brasschaat, Belgium, November
Porcupine (Erethizon dorsatum) on tundra. North Slope, Alaska, USA. September
RF- Trees reflecting in Loch Vaa, Cairngorms National Park, Scotland, UK, October 2012
Maple leaves backlit on lightbox, Broxwater, Cornwall, UK. November 2013
Melting ice on Jokulsarlon beach with moon. Southern Iceland, Europe, November 2012
Popradske Pleso, looking toward mountain range, in late evening light, with reflections and mist. Tatra Mountains, Slovakia June 2012
Trees reflecting in Loch Vaa, Cairngorms National Park, Scotland, UK, October 2012
Mixed species woodland in autumn, including Silver birch trees (Betula pendula), Peak District National Park, England, UK, October
Antelope Canyon, a slot canyon with eroded sandstone patterns, Navajo Tribal Park, Arizona, USA, December 2012
Little egret (Egretta garzetta) shaking water off wings at sunset, Lake Csaj, Pusztaszer, Hungary. May. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Dallol hot spring with salt concretions coloured by sulphur, potassium and iron, Dallol Volcano, Danakil Depression, Ethiopia, March 2015
Three hairy sea spiders (Tanystylum grossifemorum) feeding on red soft coral (Gersemia rubiformis). Browning Pass, Port Hardy, Vancouver Island. British Columbia. Canada. North East Pacific Ocean
Split image of Blue shark (Prionace glauca) close to the surface, Pico Island, Azores, Portugal, Atlantic Ocean, July
Water detail abstract. Craigengillan Estate, Ayrshire, Scotland
Monarch butterfly (Danaus plexippus) hibernating, Mariposa Monarca Special Biosphere Reserve, central Mexico, January
Looking out across sea from deserted beach, Christmas Island / Kiritimati, July 2008
Aerial view of salt pans coloured red by cyanobacteria, Lake Natron, Rift Valley, Tanzania, August 2009