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Black howler (Alouatta caraya) captive, occurs in South America
Male Mantled howler monkey (Alouatta palliata) on foot bridge, La Selva Biological Station, Costa Rica
Red howler monkey (Alouatta seniculus) howling, captive, digitally modified
Red Howler Monkey (Alouatta seniculus) hanging by tail with non-prehensile tailed Peruvian red uakari monkey (Cacajao calvus ucayalii)
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Purus red howler monkey (Alouatta puruensis) sitting in the rainforest canopy. Madre de Dios, Peru. April. Cropped
Black howler (Alouatta caraya) male and female, captive, occurs in Brazil and Paraguay. With digitally added leaf pattern
Bolivian red howler (Alouatta sara), Pampas del Yacuma Protected Area, Bolivia
Colombian red howler monkey (Alouatta seniculus) mother and baby in tree. Northern Colombia
Red howler monkey (Alouatta seniculus) howling, captive
Mantled howler monkey (Alouatta palliata) female and young in tree, La Selva Biological research station, Heredia, Costa Rica
Howler monkey (Alouatta caraya) resting, Costa Rica
Mantled howler monkey (Alouatta palliata) with bokeh, Costa Rica
Black howler monkey (Alouatta caraya) baby being carried by mother, Pantanal NP, Mato Grosso, Brazil