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Young girl in traditional dress. Mursi tribe, Mago National Park. Omo Valley, Ethiopia
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Asiatic Wild Water Buffalo (Bubalus arnee), Royal Manas National Park, Bhutan
Cape buffalo (Syncerus caffer caffer) portrait, Red-billed oxpeckers (Buphagus erythrorhynchus) removing parasites. Nakuru, Kenya
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Black faced sheep ram with twisted horns, Mull, Scotland, UK. January
Scarred face of a white rhinoceros (Ceratotherum simum) that survived an attack by poachers who illegally tranquillised her, two other rhinoceroses died from their injures
Texas Longhorn cow on high country pasture, Santa Ynez Mountains foothills, Goleta, California, USA
East Caucasian tur (Capra cylindricornis) portrait, Caucasus, Russia. May
Tibetan antelope (Pantholops hodgsonii) male, Kekexili, Qinghai, Tibetan Plateau, China, November
Bighorn sheep (Ovis canadensis) close up rams head shot, Yellowstone National Park, USA, February
Skull of male Urial sheep (Ovis vignei) on snow covered slope. Himalayas near Ulley, Ladakh, India
Gemsbok (Oryx gazella) female with two calves, Namibrand Reserve, Namib Desert, Namibia, Nambia
Female American buffalo (Bison bison) walking across grassland with newborn calf, Yellowstone National Park, Wyoming, USA. May
Snow sheep (Ovis nivicola borealis) adult with lamb, huddled against the rockface, Plateau Putorana Nature Reserve, Siberia, Russia
Alpine ibex (Capra ibex) resting in mountain pasture, Ticino, Switzerland
Black rhino (Diceros bicornis) with very long horn, Lewa Wildlife Conservancy, Laikipia, Kenya. October
Alert young Roe deer (Capreolus capreolus) buck with developing horns standing on a terraced garden lawn, Wiltshire garden, UK, October
Walia ibex (Capra walie) male with Giant lobelia (Lobelia rhynchopetalum) at around 4, 400 meters near Bwahit Pass, Simien Mountains National Park, Amhara, Ethiopia, September
Black rhinoceros (Diceros bicornis) stands in evening light on Chiefas Island in Okavango DeltaBlack rhinoceros (Diceros bicornis) stands in evening light on Chiefs Island in Okavango Delta, Botswana, . This rhino was released in the Okavango Delta as part of efforts to rebuild the rhino
White rhinoceros (Ceratotherium simum) with a long horn, Marakele National Park, South Africa
Southern lechwe (Kobus leche), two males about to fight, facing each other. Okavango Delta, Botswana
Rainbow scarab (Phanaeus vindex) male on white background. Florida, USA. Controlled conditions
Scimitar-horned oryx (Oryx dammah) captive in enclosure of Souss Massa National Park, Morocco. Extinct in the Wild
Blackbuck (Antelope cervicapra), male, Tal Chhapar Wildlife Sanctuary, Rajasthan, India
Himalayan tahr (Hemitragus jemlahicus) female. Captive, Netherlands
Highland cow in snow, Glenfeshie, Cairngorms National Park, Scotland, UK, February
Saiga antelope (Saiga tatarica) males drinking Astrakhan Steppe, Southern Russia
Rear view of Wildebeest (Connochaetes taurinus) herd crossing the Mara river, Masai-Mara game reserve, Kenya
African horned viper (Cerastes cerastes), captive
Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia
European Bison (Bison bonasus) calf and mother portrait. Mefody, first calf born to reintroduced herd in Bryansk Forest. Bryansk Forest Zapovednik, Kamchatka, Russian Far East, May
Bison (Bison bison) close up head portrait with snow on fur, Yellowstone National Park, USA, February
Horned adder (Bitis caudalis) eyes and horns, Swakopmund, Erongo, Namibia
Saiga antelope (Saiga tatarica) male, standing in grassland in morning sunlight, Betpak Dala, Kazakhstan. Critically endangered
Chamois (Rupicapra rupicapra) head portrait, Schwyz, Switzerland. November
Ibex (Capra ibex) feeding on grass, head portrait, Schwyz, Switzerland. August
Two Ibex (Capra ibex) males, standing on hillside, Lucerne, Switzerland. August
Alpine ibex (Capra ibex) male, with kid on mountain ridge in heavy winter snowfall, Valsavarenche, Gran Paradiso National Park, Italy. January
Greater one-horned rhinoceros, (Rhinoceros unicornis) walking through shallow water, Bardia National Park, Terai, Nepal
Greater one-horned rhinoceros (Rhinoceros unicornis) standing in shallow water watching a White-throated kingfisher (Halcyon smyrnensis) fly by, Bardia National Park, Terai, Nepal
Male Jackson's chameleon (Chamaeleo jacksoni) reaching out as it moves between leaves, Maui, Hawaii
Black-faced sheep (Ovis domesticus) ram, head portrait, Isle of Islay, Hebrides, Scotland, UK. April
Camargue bull (Bos taurus) running through marshland, Camargue, France. October
Axolotl (Ambystoma mexicanum) backlit showing details of gills, Mexico
Female Highland cow and calf (Bos taurus) next to hay bale, Exmoor National Park, Somerset / Devon, England. November
Female Highland cow (Bos taurus) licking its calf next to hay bale, Exmoor National Park, Somerset / Devon, England. November