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Snowy owl (Bubo scandiaca) female sitting among snow covered vegetation on ground, Canada February
Golden eagle, 4th year male, Scotland. Captive bird
Detail of Cumulonimbus cloud at sunset. Nordtirol, Austrian Alps. July
Lemur leaf frog (Agalychnis lemur) Central Caribbean foothills, Costa Rica. IUCN Red List critically endangered species
Dense rainforest surrounding estuary mouth, Corcovado National Park, Osa Peninsula, Costa Rica
Spined glassfrog (Teratohyla spinosa) Central Caribbean foothills, Costa Rica
Mountain ridge at dawn, Nordtirol, Austrian Alps, June
Trumpet / Stemless Gentian (Gentiana acaulis) Nordtirol, Austrian Alps. June
Carthusian Pink flower (Dianthus carthusianorum) detail of two stigmas, Nordtirol, Austrian Alps. June
Full moon rising between the Verpeilspitze (3430m, left) and the Watzespitze (3554m, right). These peaks are part of the Glockturmkamm, the westernmost ridge of the Otztal Alps
Round-headed Rampion (Phyteuma orbiculare) detail with out of focus Golden Hawk s?beard (Crepis aurea) in the background. Nordtirol, Austrian Alps. June
Abstract of Golden Hawk s?beard (Crepis aurea) Nordtirol, Austrian Alps. June
Astrantia (Astrantia major), Nordtirol, Austrian Alps. June
Peregrine (Falco peregrinus peregrinus) on statue at Norwich Cathedral, Norfolk, June 2013
Bewicks swans (Cygnus columbianus) small flock in flight, Gloucestershire, UK February
Gannet nesting colony on Bass Rock, UK
A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands
Starfish (Asterias rubens) below kelp in the typical scenery in Thorshofn Bay, north Iceland. North Atlantic Ocean
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification)
Lemon sole (Microstomus kitt) portrait amongst seaweeds. Gardur, south west Iceland. North Atlantic Ocean
Nudibranch (Flabellina lineata) photographed in the field aquarium, Gulen, Norway. North East Atlantic Ocean
RF - Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt
Ribbon sweetlips (Plectorhinchus polytaenia) USAT Liberty Wreck, Tulamben, Bali, Indonesia. Java Sea
RF - High magnification photo of Secretary blenny (Acanthemblemaria maria) in boulder brain coral (Colpophyllia natans). East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Grey seal (Halichoerus grypus) split level view of young investigating camera lens, Lundy Island, Devon, UK, Bristol Channel, August
California sea lion (Zalophus californianus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath Eureka oil rig, Los Angeles, California, USA, Pacific Ocean
Two Golden / Masked butterflyfish, Red Sea, Eygpt
Wolf Spiders (Pardosa sp. ), male RIGHT waving palps in courtship displayWolf Spiders (Pardosa sp.), male RIGHT waving palps in courtship display. Peak District National Park, Derbyhsire, UK. May
A portrait of a nudibranch (Eubranchus tricolor) on the seabed of a Scottish loch. Loch Carron, Ross and Comarty, Scotland, Great Britain. North East Atlantic Ocean
A blue light fluorescence photo of Jewel anemones (Corynactis viridis). The blue light source excites the fluorescent pigments in the anemones, which emit light of other colours
RF - Titan triggerfish (Balistoides viridescens) swimming over hard coral (Acropora sp.) gardens. Kri Island, Raja Ampat, West Papua, Indonesia. Dampier Strait, tropical west Pacific Ocean
Trumpet / Stemless Gentian flower (Gentiana acaulis) looking down the middle of the flower, showing five-fold symmetry. Nordtirol, Austrian Alps. June
Orchid (Arachnis flosaeris) in flower, hotel garden, Ranomafana, Madagascar, Africa. Non native species originally from Indonesia
Yellow-footed rock wallaby {Petrogale xanthopus} South Australia
Sea urchin shrimp {Periclimenes colemani} on Sea urchin {Asthenosoma varium} Indonesia Sulawesi
Two female Leaf nosed twig snakes {Langaha madagascariensis} Mantadia NP, East Madagascar
Split image of Blue shark (Prionace glauca) close to the surface, Pico Island, Azores, Portugal, Atlantic Ocean, July
Polar bears (Ursus maritimus) female with cubs aged two years travelling along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
Wild cat in pine forest {Felis silvestris} Cairngorms NP, Scotland, UK
Wide angle shot of Elephant herd taken with remote control camera during filming of BBC television programme Elephants - Spy in the Herd 2003
Solitary Leopard {Panthera pardus} walking along sand dune Namibia, Southern Africa
Australian gannets in flight {Morus serrator} Australia
Quiver tree {Aloe dichotoma} view up through canopy Namib-Naukluft NP, Namibia 2004
Eastern spinner dolphin {Stenella longirostris orientalis} W Mexico, Pacific
Tail fluke of diving Sperm whale {Physeter macrocephalus} Kaikoura, New Zealand, Pacific
Southern right whale dolphins porpoising {Lissodelphis peronii} Kaikoura, New Zealand
Melting iceberg on coast Qaanaaq, Greenland
Emperor penguin adults and chicks going to colony {Aptenodytes forsteri} Auster EP rookery, Antarctica
