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Grizzly bear (Ursus arctos horribilis) female with cub riding on back, Katmai National Park, Alaska, USA, August
Sea otter (Enhydra Lutris) with forelegs raised, Sitka Sound, Alaska, USA, August
Arctic fox (Vulpes lagopus), juvenile looking at camera, portrait, winter pelage. Dovrefjell National Park, Norway. September
Arctic fox (Vulpes lagopus) juvenile sniffing ground, winter pelage. Dovrefjell National Park, Norway. September
European hare (Lepus europaeus), Wirral, England, UK, May
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Japanese Macaques (Macaca fuscata) in hot springs, Jigokudani, Nagano Prefecture, Honshu, Japan
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Proboscis monkey (Nasalis larvatus) male, Sabah, Borneo
Sycamore moth (Acronicta aceris) caterpillar. Surrey, England, UK, September
RF - Brown hare (Lepus europaeus) adult in arable field, Scotland, UK, August
RF- North American river otter (Lutra canadensis) captive, occurs in North America
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Golden-mantled tamarin (Saguinus tripartitus) captive
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
Moutain gorilla (Gorilla beringei beringei) close up of hand, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Highland Cow (Bos taurus) close-up, Isle of Mull, Inner Hebrides, Scotland, April
Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February
Siberian musk deer (Moschus moschiferus) male with tusks grooming, captive, Midlothian deer enclosure, UK, vulnerable species
RF - Raccoon (Procyon lotor) in autumn foliage. Connecticut, USA. October
RF- European brown bear (Ursus arctos) with mouth open, Kuhmo, Finland. July
RF- Close-up of Highland cow (Bos taurus) showing thick insulating hair covering face. Isle of Lewis, Outer Hebrides, Scotland, UK, April
Tricolour merle Dachshund x Poodle Daxie doodle dog
Giant anteater (Myrmecophaga tridactyla) walking in habitat, Hato El Cedral. Llanos, Venezuela
Yellow mongoose (Cynictis penicillata) standing on hind legs with young, Kgalagadi Transfrontier Park, South Africa, February
Red Howler Monkey (Alouatta seniculus) hanging by tail with non-prehensile tailed Peruvian red uakari monkey (Cacajao calvus ucayalii)
Yellow Guinea pig standing up and squeaking, against white background
Maned wolf (Chrysocyon brachyurus) searching for food, Piaui, Cerrado, Brazil, South America
RF- Head portrait of a European grey wolf (Canis lupus), with a second in the background, captive, Norway, February
Chacma baboon (Papio hamadryas ursinus) portrait of adult male. deHoop Nature reserve. Western Cape, South Africa
Polar bear (Ursus maritimus) portrait, female. Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
RF - Red Deer stags (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Red squirrel (Sciurus vulgaris) feeding, Black Isle, Scotland, UK, February
Hazel Dormouse (Muscardinus avellanarius) hibernating amongst leaves and acorns. Captive. Leicestershire, UK, October
RF- Male Dog Otter / Eurasian Otter (Lutra lutra) head profile, captive, England
Young Japanese macaque (Macaca fuscata) kissing another macque in the thermal hot springs, Jigokudani, Japan, January
RF - Portrait of Harp seal (Phoca groenlandicus) pup on sea ice. Magdalen Islands, Gulf of St Lawrence, Quebec, Canada
Thomass Leaf Monkey / Langur (Presbytis thomasi). Gunung Leseur National Park, Sumatra, Indonesia. Endemic to Indonesia
African golden emperor moth (Gonimbrasia krucki), close up of male head, Kenya, Africa Controlled conditions
Wild cat (Felis sylvestris) in snow, Vosges, France, February
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
North American River Otter (Lontra canadensis) two swimming near the surface with autumn leaves reflected in water, Acadia National Park, Maine, USA, October
Muskox (Ovibos moschatus) in tundra with mountains in background, Dovrefjell National Park, Norway. September 2018
Okarito Brown Kiwi (Apteryx rowi) feather detail resembling mammalian hair. Okarito Forest, Westland, South Island, New Zealand
Comtois Horse (Equus caballus). Camargue reserve, Rhone, France, September
Close up of muzzle and mane of Shetland Pony, Shetland Islands, Scotland, UK
Moth caterpillar on leaf, tubercles with urticating hairs. Wuliangshan Nature Reserve, Jingdong, Yunnan Province, China
