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A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
School of black rockfish (Sebastes melanops) shelter in a Bull kelp forest (Nereocystis luetkeana). Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada
Coral mushrooms, coral banks are eroded by change in sea level, tidal current and wind, Passe Femme / Femme channel, Aldabra, Indian Ocean 2006
Giant coreopsis (coreopsis gigantea) inside the crater, Zapato Islet, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
Galapagos land iguana (Conolophus subcristatus) near the summit of Volcan La Cumbre, Fernandina Island, Galapagos
Morey eel (Muraena helena) in seagrass meadow (Posidonia oceanica) Punta Carena, Capri Island, Sorrentine Peninsula, Italy, Tyrrhenian Sea, Mediterranean
Auroras salamander (Salamandra atra aurorae), endemic to Asiago plateau, Italy, August
Mute swan (Cygnus olor) group of young swimming in a drainage channel with a parent in the background, Tealham Moor, Somerset Levels, UK, September
Langur monkey (Semnopithecus) climbing tree in misty forest at dawn, India, November
Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Nubian ibex (Capra nubiana), young females standing on rocks, Dead Sea, Israel, May
Nubian ibex (Capra nubiana), male standing in dry environment, Negev desert, Israel, April
Grey seal (Halichoerus grypus) female swims over oarweed (Laminaria digitata) in shallow water. Farne Islands, Northumberland, England, United Kingdom. North Sea
Gentoo penguin (Pygoscelis papua), gathering nesting pebbles, Beagle Channel, Patagonia, Argentina
Caribbean reef shark (Carcharhinus perezi) swimming over a coral reef, watched by a Nassau grouper (Epinephelus striatus) surrounded by common sea fans (Gorgonia ventalina)
Reef manta (Mobula alfredi) visiting a cleaning station on a coral reef with a school of Onespot snappers (Lutjanus monostigma)
Oxlip (Primula elatior) in alpine pasture following snow melt, mountains in background. Col di Rodella, Fassa Valley, Dolomites, Trentino, Italy. June 2019
Lemon shark pups (Negaprion brevirostris) in mangrove forest, Eleuthera, Bahamas
Mealy Parrots (Amazona farinosa) feeding at the wall of a clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Red-and-green Macaws (Ara chloroptera) in rainforest, Tambopata National Reserve, Peru, South America
Crested guan (Penelope purpurascens), Finca Arroyo Negro, El Triunfo Biosphere Reserve, Chiapas, southern Mexico, April
Lanzas salamander (Salamandra lanzai), endemic to Cottian Alps, Monviso massif, Italy
Polar bear (right, foreground) female and her triplets age 23-months triplets, Hudson Bay, Manitoba, Canada. November
Long-eared owl (Asio otus) looking down from a tree in autumn, Wijchen, The Netherlands, December. Second Prize Winner, Birds in their Environment Category
Kamchatka brown bear (Ursus arctos) mother and yearling cubs on autumn tundra, Lorran Peninsula, Kronotsky Nature Reserve, Kamchatka Peninsula, Russia
Zebra / leopard shark (Stegostoma fasciatum) swimming over the reef, Vaavu atoll, Maldives, Indian Ocean
Bedriagas rock lizard (Archaeolacerta bedriagae) is Sardinia, Italy. Endemic
Dwarf forest chameleon (Rhampholeon boulengeri), Nyungwe Forest National Park, Rwanda
Asp Viper (Vipera aspis) with field full of dandelions, Italy
Common poppies (Papaver rhoeas) in field, Chicklade, Wiltshire, England, July
Dense flock of Common teal (Anas crecca) with a few Wigeon (Anas penelope) and Northern shoveler (Anas clypeata) in flight over others resting on partly frozen flooded marshes
Vochysia tree and rainforest landscape, Copalinga Reserve, Ecuador, December 2018
Siberian ibex (Capra sibirica) silhouetted on ridge, Tian Shan mountains, Xinjiang, China
Split level of Sunflower stars (Pycnopodia helianthoides) on rocks underwater, Minstrel Island, British Columbia, Canada. March
Split level of Acorn barnacles (Balanus glandula) just below the surface, Seven Tree Island, Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Split-level of Cormorants (Phalacrocorax sp. ) standing on rocks with Sea lettuce (Ulva spSplit-level of Cormorants (Phalacrocorax sp.) standing on rocks with Sea lettuce (Ulva sp.) seen underwater, Hussar Bay, Nigei Island, Queen Charlotte Strait, British Columbia, Canada. September
Split level of a Lions mane Jellyfish (Cyanea capillata) and Feather-boa kelp (Egregia menziesii), Seven-tree Island, Browning Pass, Queen Charlotte Strait, British Columbia, Canada. September
Schooling rockfishes (Sebastes spp. ) with Bull kelp (Nereocystis luetkeana)Schooling rockfishes (Sebastes spp.) with Bull kelp (Nereocystis luetkeana). Different species of rockfish often swim together in mixed schools, as here
Egyptian vulture (Neophron percnopterus) adult with the Haggeher mountains in the background, Socotra Island, Yemen, March
Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists
White-cheeked pintail (Anas bahamensis) flock on water, Fernandina Island, Galapagos
Santa Fe land iguana (Conolophus pallidus) next to Tree Prickly pear (Opuntia echios) Santa Fe Island, Galapagos
Galapagos hawk (Buteo galapagensis) at nest with landscape. Cape Hammond, Fernandina Island, Galapagos
Brown pelican (Pelecanus occidentalis) landing reflected in tide pool, Puerto Egas, Santiago Island, Galapagos
Galapagos hawk (Buteo galapagoensis) perched, and landscape of habitat. Pinta Island, Galapagos
Alcedo giant tortoises (Chelonoidis vandenburghi) and volcanic landscape with fumeroles, Alcedo Volcano, Isabela Island, Galapagos
Flock of Galapagos dove (Zenaida galapagoensis) perched on rock, Santiago Highlands, Galapagos
Galapagos land iguana (Conolophus subcristatus) and landscape of Fernandina Volcano, Galapagos
