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Scarlet macaws (Ara macao) on salt lick, Tambopata, Madre de Dios, Peru
Scarlet macaws (Ara macao) and Blue and yellow macaws eating clay close to the Tambopata river. Tambopata Reserve, Madre de Dios, Peru
Greater flamingo (Phoenicopterus ruber) flock in flight, Pont de Gau, Camargue, France, May
Northern gannets (Morus bassanus) diving for fish. Bempton, Yorkshire, UK. August
Starling (Sturnus vulgaris) murmuration / mass gathering at winter roost, Salisbury Plain, Wiltshire, UK January
Chocolate-dip chromis (Chromis dimidiata) male over out of focus hard coral (Porites nodifera). Sha ab Claudia, Fury Shoal, Egypt. Red Sea
Maldive anemonefish (Amphiprion nigripes) shoal swimming in the current flowing over a Magnificent sea anemone (Heteractis magnifica). Baa Atoll, Maldives. Indian Ocean
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF - Long exposure of a school of yellowtail surgeonfish (Prionurus punctatus), these fish school to help them access the best feeding areas, which are controlled by aggressive damselfish
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Leafcutter ants (Atta sp) colony harvesting a banana leaf, Costa Rica. 3rd place in the Insects and Spiders category of the Terre Sauvage Nature Images Awards competitions 2015
School of Blackfin barracuda (Sphyraena qenie), Kimbe Bay, West New Britain, Papua New Guinea
Yellow and blueback fusilier (Caesio teres) schooling over coral reef, West Papua, Indonesia
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Yellow and blueback fusilier (Caesio teres) shoal, Similan Islands, Andaman Sea, Thailand
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
African elephant (Loxodonta africana) herd silhouetted in distance walking through a storm, Masai-Mara Game Reserve, Kenya. September
Wildebeest (Connochaetes taurinus) herd walking in a line in the rain at sunset, Masai-Mara Game Reserve, Kenya. September
Herd of Tibetan Wild Asses / Kiang (Equus kiang) ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India, October 2012
RF - Flock of Boat-tailed grackle (Quiscalus major) on wires at dawn. Crystal River, Florida, United States
Great white shark (Carcharodon carcharias) portrait, Guadalupe Island, Mexico, Pacific Ocean
A school of Southern stingrays (Hypanus americanus) on a sand bar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Group of lions (Panthera leo) at night, Masai Mara, Kenya. Photographed using Starlight Camera technology without artificial lighting
Flock of Kelp / Southern black backed Gulls (Larus dominicanus) flying over iceberg. Graham Passage, Antarctic Peninsula, Antarctica. February. (digitally altered image)
RF - Spinner dolphins (Stenella longirostris) pod swimming over a shallow sandy lagoon in a coral reef. Sataya Reef, Fury Shoal, Egypt. Red Sea
Aerial view of Humpback whales (Megaptera novaeangliae) Bay of Fundy, Canada. November
Caribbean flamingo (Phoenicopterus ruber) flying, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January
King penguins (Aptenodytes patagonicus) breeding colony with adults and juveniles, snowy mountains in background. Salisbury Plain, South Georgia. November 2018
Andean Flamingoes (Phoenicoparrus andinus) on the Altiplano, Sur Lipez Province, Potosi Department, Bolivia. December 2013
Sardine bait ball (Sardinops sagax) escaping Striped marlin (Tetrapturus audax) that is attempting to feed on them. West Coast of Baja California Peninsula, Mexico. Pacific Ocean
Caribbean flamingos (Phoenicopterus ruber) line of adults standing in water, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
Kittiwakes (Rissa tridactyla) flying over surf, Vardo, Norway March
A shoal of Golden grey mullet (Liza aurata) swim in the surf breaking on the shore of El Hierro Island. Canary Islands, Spain. East Atlantic Ocean
Greater flamingo (Phoenicopterus roseus) flock standing in lake as others fly overhead and one bird flaps its wings, Camargue, France, January
Snares crested penguin (Eudyptes robustus) colony standing on rocky shoreline behind band of washed up kelp, with a New Zealand fur seal (Arctocephalus forsteri) resting on rock beside them
Lesser flamingo (Phoeniconaias minor) flock resting at sunset. Amboseli National Park, Kenya. July
Flock of American flamingos (Phoenicopterus ruber) gathered along beach before flying to other island in search of better feeding ponds. Floreana Island, Galapagos Islands, Ecuador
Northern gannets (Morus bassanus) breeding colony nesting on sea cliffs, Hermaness, Shetland, Scotland, UK. July
Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp. ) hunting a large school of Sardines (Sardina sp)Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp.) hunting a large school of Sardines (Sardina sp.), Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Group of California sea lions (Zalophus californianus) playing near the sea surface, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp)Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp.) being followed by a California sea lion (Zalophus californianus), Espiritu Santo Island
California sea lion (Zalophus californianus) close up with group playing in shallow water behind, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Aerial view of massive arribada of Olive ridley turtle (Lepidochelys olivacea), with over 300, 000 females coming ashore to nest on 3 km of 15 km beach over period of three days and nights
