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Turnstones (Arenaria interpres) and Purple sandpipers (Calidris maritima) over wintering on the beaches of the Orkney Isles, Scotland, UK, April
Orca (Orcinus orca) surfacing, Shetland, Scotland, UK, August
Gannet (Morus bassanus) diving for fish, Noss, Shetland Islands, Scotland, UK, August
Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
White-fronted goose (Anser albifrons) and Taiga bean goose (Anser fabalis rossicus) flocks in flight and on water. Latvia. April
White pelicans (Pelecanus erythrorhynchos) at Bear River Migratory Bird Refuge with an approaching storm in May, Utah, USA, May
Wilsons Phalaropes (Phalaropus tricolor) flock flying over tufa formations at Mono Lake in late summer, with reflection, California, USA. July 2013
Avian feeding frenzy with a flock of White Pelicans (Pelecanus erythrorhynchos) swimming along a channel chasing school of fish
Mixed Blackbird flock, mostly Red-winged Blackbirds (Agelaius phoeniceus), in flight against backdrop of autumn foliage in November, Bosque Del Apache National Wildlife Refuge, New Mexico, USA
Snow buntings (Plectrophenax nivalis) flock in flight during a snowstorm, New York, USA, January
Snow Geese (Chen caerulescens) large flock in pool at Bosque Del Apache National Wildlife Refuge with setting full moon at sunrise in December, New Mexico, USA. December
Llama (Lama glama) herd, Laguna colorada. Altiplano, Bolivia
Vicuna (Vicugna vicugna), five running across salt flat with mountain in background. Salar de Ascotan, Chile. September 2018
Griffon vulture (Gyps fulvus) landing on coast next to flock. Cantabria, Spain, March
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK
Black-headed gull (Chroicocephalus ridibundus) flock flying over a frozen lake in falling snow, Wiltshire, UK, March
Greater flamingos (Phoenicopterus roseus) flock, Sado Estuary, Portugal. January
Thomsons gazelle (Eudorcas thomsonii) herd. Serengeti National Park, Serengeti, Tanzania
Piraputanga, (Brycon hilarii) reflected on the water surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June
Bohar snapper (Lutjanus bohar) school at Ras Mohammed, Shark Reef, Ras Mohammed, Sinai, Egypt. Red Sea
Masai giraffe (Giraffa camelopardalis tippelskirchi) male, females and calf standing close together, Masai Mara Game Reserve, Kenya
Whale Shark (Rhincodon typus) and Golden trevally (Gnathanodon speciosus) Cenderawasih Bay, West Papua, Indonesia
Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
Common redpoll (Acanthis flammea), flock feeding, Finland, January
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Indian wild ass (Equus hemionus khur), group standing together, Little Rann of Kutch, Gujarat, India
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, Altiplano, Bolivia
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
European Chubb (Squalius / Leuciscus cephalus) in cave, Plitvice Lakes National Park, Croatia. November
School of Munks devil ray (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Red Deer (Cervus elaphus) herd in forest in snow, Cairngorms National Park, Scotland, UK. December
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Pink-footed goose (Anser brachyrhynchus) flock flying in mist, Cromarty Firth, Highlands, Scotland, UK April
Curlew (Numenius arquata) roosting on saltmarsh, Cromarty Firth, Scotland, UK April
Greater flamingos (Phoenicopterus roseus) flock reflected, feeding in water, Pont de Gau Ornithological Park, Camargue, France, March
Leaf-cutter ants (Atta cephalotes) carrying pieces of leaf that they have harvested back to their underground fungus garden in their nest, Osa Peninsula, Costa Rica
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Bigeye trevally (Caranx sexfasciatus) forming a circular shoal in blue water, on the drop off of a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
RF - Red-and-green macaw (Ara chloropterus) flock feeding and flying. Wall of clay lick, Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
King penguin (Aptenodytes patagonicus) colony. Right Whale Bay, South Georgia. September
