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Scalloped hammerhead shark shoal (Sphyrna lewini) Wolf Island, Galapagos
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Short-finned pilot whale (Globicephala macrorhynchus) calf with fetal lines characteristic of newborn specimens, with pod. Tenerife, Canary Islands
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Small flock of Lapwings (Vanellus vanellus) flying in to rest after migration. Cresswell Pond, Druridge Bay, Northumberland, England, UK, September
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Young grey seal pup (Halichoerus grypus) recently born on a beach in Orkney, Scotland, UK, April
Orca (Orcinus orca) surfacing and blowing, Shetland, Scotland, UK, August
Common Seals (Phoca vitulina) hauled out on sand bank at Blakeney Point, Norfolk, England, UK, August
Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico
School of Cow breams (Sarpa salpa ) Azores, Atlantic ocean
Abstract interpretation of Sandhill Cranes (Grus canadensis) at roost during spring migration in March, Rowe Sanctuary, Kearney, Nebraska, USA. March. Long exposure
Huge flock of White ibis (Eudocimus albus) in breeding plumage, flying to their mangrove-covered island rookery, Tampa Bay, Florida, USA, March
Colonies of King penguins 1+Aptenodytes patagonicus+1 and Southern Elephant Seals 1+Mirounga leonina+1 on beach at St. Andrews Bay, South Georgia. November
Himalayan honey bees (Apis dorsata laboriosa) on comb, Mount Namjagbarwa, Yarlung Zangbo Grand Canyon National Park, Tibet, China. October
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Fish seeking shelter around floating tree in open ocean. Off coast of Cocos Island National Park, Costa Rica
Munks pygmy devil ray / Munks mobula (Mobula munkiana) large school from the air, Baja California, Mexico. Large group gathered possibly for mating purposes
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Flock of Tree sparrows (Passer montanus) in flight, with a Greenfinch (Chloris chloris) blurred motion, Pacsmag, Hungary
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Pilot whales (Globicephala macrorhynchus) just below surface, Tenerife, Canary Islands, Atlantic Ocean
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
Gorgonian / Sea whip coral (Ellisella ceratophyta) with Ring tailed cardinalfish (Apogon aureus) swimming in front, West Papua, Indonesia
Coral reef with soft corals (Dendronephthya sp. ), encrusting sponges and Pygmy sweepersCoral reef with soft corals (Dendronephthya sp.), encrusting sponges and Pygmy sweepers (Parapriacanthus ransonetti) Similan Islands, Andaman Sea, Thailand
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Aerial view of a herd of wild White rhinoceros (Ceratotherium simum) running free on Chiefs Island, Okavango Delta, Botswana
School of Perch (Perca fluviatilis) amongst Water lilies (Nuphar lutea) lit by the suns rays, Lake Bourget, Alps, Savoie, France, June
Springbok (Antidorcas marsupialis) herd fleeing predator, Kgalagadi Transfrontier Park, South Africa
Male Lion (Panthera leo) Romeo in habitat, from the Marsh Pride, Masai Mara, Kenya
Semipalmated sandpiper (Calidris pusilla) flock in flight over water, Johnsons Mills Reserve, Bay of Fundy, New Brunswick, Canada, July
Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project
Siberian ibex (Capra sibirica) herd, Altai Mountains, Gobi Desert, Mongolia. November
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Acrobat ant (Crematogaster sp) tending aphids on white oak (Quercus sp) Washington State Park, Pennsylvania, USA September
Lesser flamingo (Phoeniconaias minor) flock, Bogoria Game Reserve, Kenya
Caribbean Flamingo (Phoenicopterus ruber) chick group walking around the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Aerial view of Caribbean Flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May
Flock of White ibis (Eudocimus albus) in breeding plumage, at rookery on water sFlock of White ibis (Eudocimus albus) in breeding plumage, at rookery on waters edge of the mangrove-covered island, Tampa Bay, Florida, USA, March
Meerkats (Suricata suricatta) standing alert, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
Snorkeller with shoal of Piraputanga (Brycon hilarii), Rio Olha d Agua at Cabeceira do Prata, Bonito, Mato Grosso do Sul, Brazil
Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June