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Shaugh Bridge in early spring, Shaugh Prior, near Plympton, Dartmoor National Park, Devon, UK. May 2018
Hawes, Wensleydale, Yorkshire Dales National Park, North Yorkshire, England, UK. June 2017
Northern pike (Esox lucius) amongst Shining pondweed (Potamogeton lucens). Lake Neuchatel, Switzerland, September. Photographed for The Freshwater Project
Mute swan (Cygnus olor) adult with cygnets on water, Woodwalton Fen, Cambridgeshire, UK, June
Ghost glass frog (Centrolenella ilex) portrait, Costa Rica
Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica
Hornbeam (Carpinus betulus) leaves in spring. Dorset, UK May
Indian peafowl (Pavo cristatus) peacock displaying feathers, captive, occurs in South Asia
European roller (Coracias garrulus) in flight, Pusztaszer, Hungary, May
Splendid leaf frog (Cruziohyla calcarifer) La Selva Field Station, Costa Rica
Rainbow river, Rainbow Springs State Park, Florida, USA. January. Photographed for The Freshwater Project
Golitha Falls, River Fowey flowing through wooded valley with moss covered trees, Nr St Cleer, Cornwall, UK, May 2012
White-lipped green tree frog (Litoria infrafrenata) clinging on to a branch, Etty Bay, Wet Tropics World Heritage area, Queensland, Australia
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Kakapo (Strigops habroptilus) close up showing sensory facial feathers, Codfish Island / Whenua Hou, Southland, New Zealand, February. Critically endangered
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise
Pine marten (Martes martes) portrait, Ardnamurchan Peninsula, Scotland, August. Highly commended in the Portrait category of the BWPA (British Wildlife Photography Awards) Competition 2016
Sand lizard male {Lacerta agilis} Ainsdale NNR, Lancashire, UK
Chlorophyte or green alga (Volvox aureus) in pond water. UK. The larger spheres are approximately 0.5mm in diameter. Digital composite image
Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013
Dolgoch Falls, Gwynedd, Wales, UK, May 2015
Seven Little Bee-eaters (Merops pusillus) in a row, one facing away, Masai Mara, Kenya
Musk lorikeet (Glossopsitta concinna) feeding on flowers in a Eucalyptus tree, south-east Tasmania, Australia, February
Rice (Oryza sativa) terraces. Jatiluwih Green Land, Bali, Indonesia. 2015
Palms growing in front of Rice (Oryza sativa) terrace. Jatiluwih Green Land, Bali, Indonesia. 2015
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Candover Brook, tributary of the River Itchen, chalk stream Hampshire, England, UK, June. Photographed for The Freshwater Project
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) Mindo, Pichincha, Ecuador
Ewens Ponds, spring fed limestone ponds at the Ewens Ponds Conservation Park, South Australia, Australia March 2015. Photographed for The Freshwater Project
Irish damselfly (Coenagrion lunulatum) on leaf, Montiaghs Moss, County Antrim, Northern Ireland, UK, June
Looking up through carpet of Bluebells (Endymion nonscriptus) to Beech (Fagus sylvatica) woodland canopy, Micheldever Woods, Hampshire, England, UK, April
Himalayan monal (Lophophorus impejanus) male perched on branch, Uttarakhand, India
Green hairstreak butterfly (Callophrys rubi) on hawthorn leaf, Wiltshire, England, UK, April
Madagascar green-emerald giant pill millipedes (Zoosphaerium neptunus) Andasibe-Mantadia National Park. Madagascar
Resplendent quetzal (Pharomachrus mocinno), captive, Chiapas, southern Mexico, IUCN Near Threatened, April
Mother of thousands (Kalanchoe daigremontiana), viviparous growth with vegetative propagation of plantlets on leaf margins. In cultivation
Dry-stone walls and barns in Wensleydale, Yorkshire Dales National Park, North Yorkshire, England, UK, June 2016
American green tree frog {Hyla cinerea}
Red eyed tree frog (Agalychnis callidryas) portrait Captive
Leaves of a Ginkgo tree or Maidenhair tree (Ginkgo biloba) Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Reticulated glass frogs (Hyalinobatrachium valerioi) pair in amplexus, Osa Peninsula, Costa Rica
Underwater shot of aquatic plants in an oxbow lake of the River Aare, Haftli nature Reserve, Buren an der Aare, Switzerland
Senegal chameleon (Chamaeleo senegalensis) walking over flat ground, Allahein river, The Gambia
Watercolour painting of Common hawthorn (Crataegus monogyna) berries. Botanical illustration by Linda Pitkin
Cotton or hibiscus harlequin bug (Tectocoris diophthalmus) nymph feeding on a green leaf, Brisbane, Queensland, Australia. April
Lantern bug (Pyrops whiteheadi), Danum Valley, Sabah, Borneo
Achnambeithach Cottage on the shores of Loch Achtriochtan at the foot of Aonach Dubh, Glencoe, Lochaber, Highland, Scottish Highlands, Scotland, UK. June 2018
Lake Paradise, Marsabit National Park, Kenya
Large leaf of Giant Elephants Ear (Alocasia odora), in rainforest, Yangminshan, Taiwan
Western hemlock (Tsuga heterophylla) in Douglas-fir forest (Pseudotsuga menziesii), Oregon, USA, June
Lord Howe Island stick insect (Dryococelus australis), Bristol Zoo Gardens, Bristol, UK
Sambava Panther Chameleon (Furcifer pardalis), Eastern Madagascer, controlled conditions
Yellow star of Bethlehem (Gagea lutea) in woodlands of th Schonbrunn Palace in Vienna, Austria, March
Beech (Fagus sylvatica) forest, view into canopy from below. Peerdsbos, Brasschaat, Belgium. April 2019
Sumatran pit viper (Trimeresurus sumatranus) juvenile on tree trunk, in tropical rainforest. Danum Valley, Sabah, Borneo, Malaysia
Sesame (Sesamum indicum), green unripe pods. Sichuan Province, China
Amazon rainforest canopy view with flowering Bromeliad epiphytes growing on a branch of a giant Cieba tree. Tiputini Biodiversity Station, Amazon Rainforest, Ecuador, January
Laurisilva Forest, trees covered in lichen / moss at Terceira Island, Azores, August 2011
Nitsches bush Viper (Atheris nitschei) captive, from Uganda and Congo
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) in flight, about to land. Ecuador, May
Water lilies (Nymphaea alba) lit by the suns rays, seen under water. Alps, Ain, France. June. Highly commended in the the Underwater World category
English oak (Quercus robur) ripening acorns, Dorset, UK September
Yew berries (Taxus baccata) Berwickshire, Scotland, September
Stunted oak woodland covered in moss, Wistmans Wood, Devon, UK. August. Highly Commended in the Wild Woods category of the British Wildlife Photography Awards (BWPA) competition 2013
Damaged forest overgrown by various vines, a typical scene in western part of Dominica, West Indies. After hurricane Maria an unprecedented number of vines took advantage of suddenly open spaces in
Larvae of Birch sawfly (Craesus septentrionalis) in alarm position, Upper Bavaria, Germany. October
Australian green tree frog (Litoria caerulea) camouflaged on Palm leaf. Lake Argyle, Kununurra, Western Australia
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Satiny Parrot snake (Leptophis depressirostris) Santa Rita, Costa Rica
Coral (Fungia sp) fluorescence at night, captured using very powerful electronic strobes. This image is a comparison with image 8118530, Red Sea
Aerial view of shifting sand banks off the coast of the Bazaruto Archipelago, Mozambique, May 2011
RF- Aerial view of the Okavango delta, Botswana, May 2010
Fossil ammonite viewed under UV light, Desmoceras spp. upper early cretaceous, Albian stage, Mahajanga province, Madagascar
Moss covered fallen trees in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
Seedling cone plants (Conophytum subfenestratum), approximately two months after sowing. Focus-stacked image
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May
Aurora Borealis over Vestrahorn, Stokksnes, Hofn, Iceland. March 2018
Chinese moon moth (Actias dubernardi, female at rest amongst leaves. Dayaoshan, Jinxin, Guangxi, China
Marsh fritillary (Euphydryas aurinia) on fern. South of Casteil, Pyrenees Orientales, south west France. May
Oleander Hawkmoth (Daphnis nerii) resting, Southern Sicily, Italy. March
American moon moth (Actias luna) male, North America