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Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Green turtle (Chelonia mydas) swimming near mangroves, Elizabeth Bay, Isabela Island, Galapagos
Male Blue-footed booby (Sula nebouxii) walking on sand, San Cristobal Island, Galapagos, South America
Vampire ground finch (Geospiza septentrionalis) attacking large chick to drink blood, after it was wounded by aggressive adult booby, Wolf Island, Galapagos
Pinzon giant tortoise (Chelonoidis duncanensis), saddleback type typical of arid island, their long necks and raised shell allowing them to browse on cacti
Spotted eagle ray (Aetobatus narinari) Kicker Rock, San Cristobal Island, Galapagos
Galapagos penguins (Spheniscus mendiculus) underwater with shoal of fish, Tagus Cove, Isabela Island, Galapagos
Floreana giant tortoise hybrid descendant (Chelonoidis elephantopus), Fausto Llerena Giant Trotoise Breeding Centre, Galapagos National park Directorate, Puerto Ayora, Santa Cruz Island, Galapagos
Scalloped hammerhead shark shoal (Sphyrna lewini) Wolf Island, Galapagos
Sally lightfoot crabs (Grapsus grapsus) on rock with waves, Cape Douglas, Fernandina Island, Galapagos
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Green turtle (Chelonia mydas) female laying eggs, North coast, Santa Cruz Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Galapagos sea lion (Zalophus wollebaeki) pup on shore, Galapagos
Alcedo giant tortoises (Chelonoidis vandenburghi) group resting in water to keep cool, Alcedo Volcano, Isabela Island, Galapagos
Western Santa Cruz giant tortoise (Chelonoidis porteri) portrait, Highlands, Santa Cruz Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) males in breeding colouration on rocks, with sun halo, Floreana Island, Galapagos
Green turtle (Chelonia mydas), female returning to the sea after egg laying, Puerto Pajas, Isabela Island, Galapagos
Remoras (Remora sp. ) hitchhiking on the tail of a large (13-15m) female whale sharkRemoras (Remora sp.) hitchhiking on the tail of a large (13-15m) female whale shark (Rhincodon typus) swims in open water. Wolf Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Remoras (Remora sp. ) hitchhiking on the tail of a large (13-15mRemoras (Remora sp.) hitchhiking on the tail of a large (13-15m) female whale shark (Rhincodon typus) swims in open water. Wolf Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
American flamingo (Phoenicopterus ruber) pair in courtship, group of three feeding, Punta Cormorant, Floreana Island, Galapagos
Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos
Galapagos penguin (Spheniscus mendiculus) incubating egg, Puerto Pajas, Isabela Island, Galapagos
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) and Sally-lightfoot crabs (Grapsus grapsus) on rocks at high-tide with sea spray, Galapagos Islands, South America
Waved albatross (Phoebastria irrorata) head portrait, Espanola Island, Galapagos, South America
Scalesia pedunculata forest, Los Gemelos, Highlands, Santa Cruz Island, Galapagos, South America