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Four Wisent / Bison (Bison bonasus) standing in a row. Bialowieza Forest, Bialowieza National Park, Poland, November
Crabeater Seal (Lobodon carcinophaga) hauled out on ice. Pleneau Island, Antarctica, January
Domestic dog, Mongrel, studio portrait
Wolverine (Gulo gulo) climbing birch tree, Kamchatka, Far East Russia, April
Moose (Alces alces) head portrait of bull standing in forest clearing, Cap Breton Highlands National Park, Nova Scotia, Canada, September
King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia
Meerkat / Suricate family group (Suricatta suricata) standing alert together, Kalahari Meerkat Project, Van Zylsrus, Northern Cape, South Africa
Big close up of the eyes of a European Brown bear (Ursus arctos) captive
Young Japanese macaque (Macaca fuscata) kissing another macque in the thermal hot springs, Jigokudani, Japan, January
RF - Portrait of Harp seal (Phoca groenlandicus) pup on sea ice. Magdalen Islands, Gulf of St Lawrence, Quebec, Canada
RF- Young male Bengal tiger, head portrait in profile (Panthera tigris tigris). Bandhavgarh National Park, India
Wild cat (Felis sylvestris) in snow, Vosges, France, February
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
North American River Otter (Lontra canadensis) two swimming near the surface with autumn leaves reflected in water, Acadia National Park, Maine, USA, October
Orang utan (Pongo pygmaeus) baby sleeping in the arms of an adult, Semengoh Nature reserve, Sarawak, Borneo, Malaysia, Endangered
Muskox (Ovibos moschatus) in tundra with mountains in background, Dovrefjell National Park, Norway. September 2018
Jaguar (Panthera onca) portrait, lying down, captive
Juvenile Snow leopard {Panthera uncia} captive
Eastern Gray Squirrel (Sciurus carolinensis) feeding in snow, Acadia National Park, Maine, USA, February
North American river otter (Lontra canadensis) two in water with autumnal trees reflected in the water, Acadia National Park, Maine, USA, October
Polar bear (Ursus maritimus) head portrait of adult scenting the wind, Arctic National Wildlife Refuge, Alaska
RF- Northern Raccoon (Procyon lotor) drinking from wetland lake. Fennessey Ranch, Refugio, Coastal Bend, Texas Coast, USA
Blue merle Collie x Poodle Cadoodle puppy looking up
Sea otter (Enhydra lutris) floating on its back at surface among kelp, Alaska, USA. Gulf of Alaska, Pacific ocean
Chocolate blue eyed Mini American Shepherd puppy
Bolonka Zwetna, puppy age 3 months, portrait against white background
Musk ox (Ovibos moschatus) two covered in snow, Wrangel Island, Far Eastern Russia, September
Samoyed dog running in in snow, Ledyard, Connecticut, USA
Grey squirrel (Sciurus carolinensis) adult and baby, against white background
Pine marten (Martes martes) juvenile male sitting on branch of a dead tree outside an old black woodpecker (Dryocopus martius) nest hole. Molde, Central Norway, September
Himalayan Blue sheep (Pseudois nayaur) Hemas National Park, Ladakh, India
Red Foxes (Vulpes vulpes) young play fighting. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Red Fox (Vulpes vulpes) curled up, with an eye open. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
European Bison (Bison bonasus) calf and mother portrait. Mefody, first calf born to reintroduced herd in Bryansk Forest. Bryansk Forest Zapovednik, Kamchatka, Russian Far East, May
European Brown Bear (Ursus arctos arctos), a large male taking a rest in a cloud of mosquitoes. Kajaani, Finland. June
Himalayan marmot (Marmota himalayana) sun bathing on rock, Mount Kailash, Tibet
Proboscis monkey (Nasalis larvatus), Sabah Malaysia, Borneo
Unau / two-toed sloth (Choloepus didactylus) portrait, French Guiana
Brown Hare (Lepus europaeus) close-up of fur and eye. Derbyshire, UK, March
Water vole (Arvicola amphibius) feeding at edge of water, Kent, UK, December
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Water vole (Arvicola amphibius) standing on hind legs sniffing blackberry, Kent, UK, September
Portrait of European river otter {Lutra lutra} captive, UK
RF- Red squirrel in autumn (Sciurus vulgaris). Strathspey, Scotland, UK
Poitou donkey (Equus africanus asinus) with shaggy coat on the island Ile de Re, Charente-Maritime, France
Polar bear (Ursus maritimus) underwater view swimming in a pool, Anchorage Zoo, Alaska, USA. No release available
Black bear (Ursus americanus) young cub portrait, Yellowstone National Park, Wyoming, USA
Brown Bear (Ursus arctos) portrait in water. Finland, Europe, June