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Female Grizzly bear (Ursus arctos horribilis) standing up, with two cubs nearby, Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Lesser flamingo (Phoeniconaias minor) portrait, Lake Bogoria, Rift valley, Kenya, Africa
Fire salamander (Salamandra salamandra) female almost ready to give birth to her larvae into the stream, Apennines. Antola Regional Park, Italy
Rangimairewhenua or Blue Lake, Nelson Lakes National Park, Southern Alps, New Zealand, February 2013. Photographed for The Freshwater Project
Great Fen / Raft spider (Dolomedes plantarius), adult female eating an invasive species of fish, Western mosquitofish, (Gambusia affinis), Alessandria, Italy, August
Snapping turtle (Chelydra serpentina) with Painted turtle (Chrysemys picta) feeding on algae on the back of the snapper, Maryland, USA. August
Split level view of blue green waters of Verzasca River flowing over granite rocks at Lavertezzo, Canton Tessin, Switzerland, September. Taken for the Freshwater Project
Olm (Proteus anguinus) close up of front limb, these only have three toes whilst the rear limbs have only two. Captive, Slovenia
Green hydra (Hydra viridissima) found in a garden pond. Derbyshire, UK. September. Winner of the Hidden Britain Category of the British Wildlife Photographer of the Year Awards (BWPA) Competition 2017
Water vole (Arvicola amphibius) feeding, Kent, UK May
RF - Female emperor dragonfly (Anax imperator) laying eggs on garden pond, Broxwater, Cornwall, UK. July
Water vole (Arvicola amphibius) feeding on aquatic plant, Kent, UK May
Great crested newt (Triturus cristatus) male in breeding condition, amongst Soft hornwort (Ceratophyllum submersum), Herefordshire, England, UK, May. Captive
Red fox (Vulpes vulpes) at edge of stream, Kent, UK. January 2013
Palmate newt (Triturus helveticus) male and female underwater, captive
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
Mallard {Anas platyrhynchos} looking down from the water surface, view from underwater, Wales, UK, 2006
Lightning storm over Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May 2012
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia
River Plym flowing fast through Dewerstone Wood, Shaugh Prior, Dartmoor National Park, Devon, England, UK. May 2015
The Northern Lights / Aurora Borealis over a river, Vik, Iceland. February 2014
Edible frog (Rana esculenta) in pond weed, Vosges, France, June
Lesser flamingo flock (Phoeniconaias) flying over lake with green algae, aerial view, Rift valley, Kenya
Florida Red-bellied Turtle (Pseudemys nelsoni) Rainbow Springs, Rainbow Springs State Park, Florida, USA, March. Taken for the Freshwater Project
Cape buffalo (Syncerus caffer) crossing the Chobe River, followed by swarm of flies, Chobe Game Reserve, Botswana, Africa
African forest elephant (Loxodonta cyclotis) in water, Lekoli River, Republic of Congo (Congo-Brazzaville), Africa. Vulnerable species
Water chestnut (Trapa natans) viewed from underwater, in Danube Delta, Romania, June
Ruffe (Gymnocephalus cernua) shoal, Danube Delta, Romania, June
European white waterlilies (Nymphaea alba) viewed from underwater in a tributary of Danube Delta, Romania, June
White water lilies (Nymphaea alba) on surface of Danube delta rewilding area, Romania
Morelets crocodile / Central American crocodile (Crocodylus moreletii) hiding in the plants at the surface of the cenote, Car Wash Cenote/ Aktun Ha, Yucatan peninsula, Mexico
Suwanee freshwater turtle (Pseudemys concinna suwanniensis) swimming beneath trees in a freshwater spring. Devil Spring, Ginnie Springs, High Springs, Florida, USA
Waterfall at mouth of Kitum cave Mt. Elgon National park, Kenya, August 2017
Underwater landscape within a mountain stream, showing water turbulence, Snowdonia NP, Gwynedd, Wales, UK, December
The Fairy Pools, on the Allt Coir a Mhadaidh river running down from the Cullin Hills, Glen Brittle, Isle of Skye, Inner Hebrides, Scotland, UK, February 2010
Young Florida manatee (Trichechus manatus latirostrus) portrait, Three Sisters Spring, Crystal River, Florida, USA. February 2010
The San Rafael waterfall, the biggest falls in Ecuador, located on the boundary of the Amazon with The Andes. San Rafael, Napo, Ecuador. February 2016
Las Golondrinas Waterfalls, Montes Azules Biosphere Reserve, Chiapas. Mexico, March 2017
Misol-ha Waterfall, Ejido San Miguel, Salto de Agua Municipality, Chiapas. Mexico, March 2017
Oxbow lake adjacent to the Karawari River, Papua New Guinea, August
Aerial view of the River Kinabatangan and riverine tropical rainforest, Sabah, Malaysia, Borneo, Asia
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
Ashness launch/jetty, Ashness, sunset, Derwent Water, The Lake District, Cumbria, UK. October 2016
Pen yr Ole Wen reflected in Llyn Idwal in the Glyderau mountains in Snowdonia, North Wales, UK, December 2016
