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Blackbird (Turdus merula) male eating winfall apples in snow, Broxwater, Cornwall, UK. December
Marsh fritillary butterfly (Euphydryas aurinia) nectaring on a Meadow buttercup (Ranunculus acris) flower in a chalk grassland meadow, Wiltshire, UK, June
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
Harvest mouse (Micromys minutus) climbing among Cow Parsley, Hertfordshire, England, UK, May, Controlled conditions
Water Vole (Arvicola amphibius / terrestris) foraging by water. Kent, UK, August
Green Woodpecker (Picus viridis) female among wind-fallen apples. Hertfordshire, England, UK, January
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Grandidiers / Giant Striped Mongoose (Galidictis grandidieri) foraging at night. Western edge of the Mahafaly Plateau, Tsimanampetsotsa National Park, south west Madagascar
Stewart Island Brown Kiwi / Southern Tokoeka (Apteryx australis lawryi), female probing by scent for sand hoppers in rotting kelp beach wrack, Ocean Beach, Stewart Island, New Zealand
Green sandpiper (Tringa ochropus) with small worm it has caught in shallow freshwater lake, Gloucestershire, UK, August
American avocet (Recurvirostra americana), adult in breeding plumage foraging by sweeping its bill from side to side through shallow water, Orange County, California, USA, April
California Gull (Larus californicus), foraging for alkali flies (Ephydra hians) by running through swarm snapping its bill, on the shore at Mono Lake, California, USA. Digital frame expansion
Tricolored heron (Egretta tricolor) adult in breeding plumage capturing fish by half-running half-flying across waters surface and stabbing with its bill, Viera Wetlands, Brevard County, Florida
Pine marten (Martes martes) foraging in Pine (Pinus sp) woodland at night. Glenfeshie, Cairngorms National Park, Scotland, UK
Eurasian badger (Meles meles), two foraging in Pine (Pinus sp) woodland. Glenfeshie, Cairngorms National Park, Scotland, UK
Badger (Meles meles) eating apples in urban garden. Sheffield, England, UK. October
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Alpha male Timber wolf (Canis lupus lycaon) hunting in snow, Hudson Bay, Manitoba, Canada
Female Red fox (Vulpes vulpes) foraging inside a rotting tree trunk, Verteskozma, Hungary. Hungary Nature Photographer of the Year Competition 2021 Highly Commended - Mammals Behaviour category
Great white egret (Ardea alba) fishing in shallow pond, Valkenhorst Nature Reserve, The Netherlands, August
Two Black swans (Cygnus atratus) on water, Noirmoutier Island, Vendee, France, July
Wild boar (Sus scrofa) big male foraging in autumn forest during the hunting season in the Ardennes, Belgium. November
Eurasian oystercatcher (Haematopus ostralegus) walking over tidal mudflats in low winter sunshine. Holy Island (Lindisfarne) tidal mudflats, Northumberland, England, UK. December
Sloth bear (Melursus ursinus) male foraging in forest understory. Panna Tiger Reserve, Madhya Pradesh, Central India
Sulawesi or Celebes c black macaque (Macaca nigra) foraging on exposed beach at low tide. Tangkoko National Park, Sulawesi, Indonesia
Eurasian jay (Garrulus glandarius) foraging in autumn leaves. London, England, UK. November
Honey bee (Apis mellifera) nectaring on a Crab apple (Malus sylvestnis) flower in a garden, Wiltshire, UK, April
Hazel Grouse (Tetrastes bonasia), Kuusamo, Finland, December
Great spotted woodpecker (Dendrocopos major) and Grey headed woodpecker (Picus canus) on tree trunk, Germany. December
Bearded Pigs (Sus barbatus) foraging on forest floor. Near Nepenthes Field Camp, mid-altitude montane forest, Sabahs Lost World, Borneo
Honey bee (Apis mellifera) with full pollen baskets feeding on Balkan anemone (Anemone blanda). In garden. Surrey, England, UK. April
Honey bee (Apis mellifera) extending proboscis to nectar on Common rue (Ruta graveolens). Green aphid (Aphidoidea) beneath flower. Surrey, England, UK. July
Cambridge vagrant butterfly (Nepheronia thalassina) nectaring on Lantana (Lantana camara). Flowers turn from yellow to pink when no nectar is provided. In garden, Tanzania. Native to tropical Americas
Honey bee (Apis mellifera) nectaring on freshly opened yellow Lantana (Lantana camara) flowers. Flowers turn pink when nectar no longer provided. In garden, Tanzania. Native to tropical Americas
Bumblebee (Bombus sp) queen feeding on Crocus (Crocus korolkowii), covered in pollen. Ansob Pass, Tajikistan. April
Bumblebee scarab beetles (Pygopleurus humeralis) feeding on Phelypaea flower (Phelypaea coccinea). Flower has marks that mimic beetle elytra wings to attract pollinators
Red bowled anemone (Anemone bucharica) with dark stamens and patterns on petal bases to attract Scarab beetle (Eulasia sp) pollinators
Coquerels sifaka (Propithecus coquereli) searching fallen fruit on forest floor, Mangatsa, Madagascar
Red squirrel (Sciurus vulgaris) foraging in forest, climbing down tree trunk, Cairngorms National Park, Scotland, October
Quinling Golden snub nosed monkey (Rhinopitecus roxellana qinlingensis) family group foraging along a small creek in a gullly. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
Grey squirrel (Sciurus carolinensis) foraging in an oak woodland. Perthshire, Scotland, Nov
Goldfinch (Carduelis carduelis) perched on Hawthorn, Cheshire, UK, October
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Pale fork-marked lemur (Phaner pallescens) in search for food at night, Kirindy Deciduous Forest, Madagascar. Lenses for Conservation project
Eurasian otter (Lutra lutra) feeding whilst on log in woodland burn, Glenfeshie, Cairngorms National Park, Scotland, UK, September
Rock pipit (Anthus petrosus) foraging for invertebrates among seaweed on the strand line as the tide rises at sunset, Cornwall, UK, September
Badger (Meles meles) walking along log at night, Mid Devon, England, September
Three 10 month old Badgers {Meles meles} browsing in leaf litter at night, Derbyshire, UK
Badger (Meles meles) standing at base of tree at night, Mid Devon, England, August
Pileated woodpecker (Dryocopus pileatus) male excavating in search of food, New York, USA. Digital composite
Pinyon Jay (Gymnorhinus cyanocephalus), gathering Pinyon Pine seeds in autumn. Mono Lake Basin, California, USA, October
Wood sandpiper (Tringa glareola) in reed bed, Finland, May
Black necked grebe (Podiceps nigricollis) adult foraging at night and catching mosquitoes. The Netherlands.March 2014. Highly commended in Asferico Awards 2017
Mint moth (Pyrausta aurata) feeding on Wild marjoram flower (Origanum vulgare), chalk grassland meadow, Wiltshire, UK, July
Black skimmer (Rynchops niger) foraging along surf line. Tierra Verde, Florida, USA
Gloucester old spot domestic pigs (Sus scrofa domestica) ears covering eyes, freerange in field, UK
Flock of American flamingos (Phoenicopterus ruber) gathered along beach before flying to other island in search of better feeding ponds. Floreana Island, Galapagos Islands, Ecuador
White-vented storm petrel (Oceanites gracilis) feeding at sea surface, in flight. Galapagos Islands, Ecuador
Marine iguana (Amblyrhynchus cristatus) emerging from sea after feeding on algae. Fernandina Island, Galapagos Islands, Ecuador
Water rail (Rallus aquaticus) wading through water, Dumfries and Galloway, Scotland, UK
Water rail (Rallus aquaticus) foraging near edge of stream, Dumfries and Galloway, Scotland, UK
Red rock crab (Grapsus adscensionis) grazing on algae from rock, Tenerife, Canary Islands, October
A pair of Flamingo tongue cowries (Cyphoma gibbosum) on a common sea fan (Gorgonia ventalina) which they predate upon, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Aerial view of Lesser flamingos (Phoeniconaias minor) foraging at dusk. Lake Magadi, Great rift valley, Kenya
Common murre / guillemot (Uria aalge) and Fulmar (Fulmarus glacialis) eggs foraged from Skoruvikurbjarg cliffs, Langanes Peninsula, Iceland. May 2018
Black guillemot (Cepphus grylle) holding fish in beak, flying back to nest, Portpatrick, Dumfries and Galloway, Scotland. June
Male Baltimore oriole (Icterus galbula) foraging in flowering Crabapple (Malus sp)Male Baltimore oriole (Icterus galbula) foraging in flowering Crabapple (Malus sp.) tree in spring, Ithaca, New York, USA. May
Komodo dragon (Varanus komodoensis) male, foraging in mangove / inter-tidal zone at low tide, Komodo Island, Komodo National Park, Indonesia. Endangered
Ocellated eagle ray (Aetobatus ocellatus) searching for food in the sand over the seafloor, Mu Koh Similan National Park, Andaman Sea, Phang-nga, Thailand
Quinling Golden snub nosed monkey (Rhinopitecus roxellana qinlingensis), family group foraging along a small creek in a gullly. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China
Giant anteater (Myrmecophaga tridactyla) foraging for ant and termite nests across savannah grassland, Caiman Ecological Refuge, Pantanal, Brazil
Black stork (Ciconia nigra) foraging in shallow water, Pusztaszer reserve, Hungary. May
Female Black woodpecker (Dryocopus martius), pecking at tree trunk, Finland. September
Spotted nutcracker (Nucifraga caryocatactes) searching for seeds on forest floor, Finland. October
Silhouette of Dunlin (Calidris alpina) foraging in shallow waters in front of the rising sun, Gulf of Gdansk, Poland. September
Darwins woodpecker finch (Cactospiza pallida) using twig as tool to extract beetle grub from dead branch, Santa Cruz Island, Galapagos, South America
Wild boar (Sus scrofa) crossing drying pond with Great white egret (Ardea alba) fishing in background, Valkenhorst Nature Reserve, The Netherlands, August
Cormorant (Phalacrocorax carbo) swimming on pond, Valkenhorst Nature Reserve, The Netherlands, August
Three Spoonbills (Platalea leucorodia) feeding in shallow pond, Valkenhorst Nature Reserve, The Netherlands, August
Mixed group of Grey heron (Ardea cinerea), Great white egret (Ardea alba) and Spoonbill (Platalea leucorodia) on misty morning in shallow pond, Valkenhorst Nature Reserve, The Netherlands, August
Black-tailed godwit (Limosa limosa) foraging, Vendeen Marsh, Vendee, France, May
Sanderling (Calidris alba) Britanny, France, October
Red knots (Calidris canutus) in winter plumage feeding co-operatively on tidal mudflats. Holy Island (Lindisfarne), Northumberland, England, UK December
Sulawesi black macaques (Macaca nigra) males foraging on exposed beach at low tide. Tangkoko National Park, Sulawesi, Indonesia