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King penguins (Aptenodytes patagonicus) crossing water to reach breeding site, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October
Elephant (Loxodonta africana) herd walking in a line, Etosha National Park, Namibia
Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Dromedary camel (Camelus dromedarius) train crossing Merzouga dunes, Tafilalt, Sahara, Morocco, March 2007
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
Mute swan (cygnus olor) family, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June
Group of California sea lions (Zalophus californianus) in a kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean
Canada goose (Branta canadensis) adult swimming leading five goslings, Ithaca, New York, USA
Female Capybara (Hydrochoerus hydrochaeris) swimming in line with young after escaping a Jaguar attack (Panthera onca palustris) in a lagoon off the Paraguay River
Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Wildebeest (Connchaetes taurinus) running down bank of the Mara River during migration, Masai Mara, Kenya
A large herd of Tien Shan Argali (Ovis ammon karelini) making tracks through snow. Naryn National Park, Kyrgyzstan, Central Asia, November
Two Common ravens (Corvus corax) in flight through snow, Finland, April
Aylesbury ducks following in a line on village green, Weedon, Buckinghamshire, UK, October
African elephant {Loxodonta africana} bulls walking in line, Etosha national park, Namibia
Family of African elephants (Loxodonta africana), Masai Mara National Reserve, Kenya, Africa
Mute swan (Cygnus olor) parent and cygnets swimming. London, UK. May
Group of Wild Boar (Sus Scrofa) trotting through forest of Rambouillet, near Paris, France. Autumn
Lion (Panthera leo) older cub playing with a young one and biting its tail, Masai-Mara Game Reserve, Kenya. Vulnerable species
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd walking, Masai-Mara Game Reserve, Kenya. October
Wild Mustang horses running, Sand Wash Basin Herd Area, Colorado, USA
Roseate Spoonbills (Platalea ajaja) in flight, Pantanal, Mato Grosso do Sul, Brazil
Burchells zebra (Equus quagga burchellii) walking in single file, Ngorongoro Crater, Tanzania
Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
Common Raven (Corvus corax) two in flight during snow storm, Kuusamo, Finland, April
Blacktip reef sharks (Carcharhinus melanopterus) swimming in shallow crystal clear water, Aldabra Atoll, Seychelles, Indian Ocean
Little Auks (Alle alle) flying low above surface in front of iceberg. Spitsbergen, Svalbard, Norway, July 2010. Highly commended in the Birds category in the GDT competition 2011
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK
Female Cheetah (Acinonyx jubatus) with juveniles crossing grassland in rain storm, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Female polar bear (Ursus maritimus) with two cubs walking in a line across snow, Churchill, Canada. November
Whale shark (Rhincodon typus), an endangered species, feeding on zooplankton in nutrient-rich murky water, being followed by a school of Cobia (Rachycentron canadum), Adang-Rawi Archipelago, Satun
Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Juvenile Mute swan (Cygnus olor) in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Female Snow leopard (Panthera uncia) with her three month cub, BioParc Doue la Fontaine, France. Captive
Elephant (Loxodonta africana) three calves walking down to Chobe River, Etosha National Park, Namibia, March
Polar bear (Ursus maritimus) with cub walking across ice field, Wrangel Island, Far Eastern Russia, March
Polar bear (Ursus maritimus) mother with cubs walking through snow, Wrangel Island, Far Eastern Russia, March
RF - Mallard (Anas platyrhynchos) female swimming with with young chicks following, Valkenhorst Nature Reserve, The Netherlands June
Four Mute swans (Cygnus olor) walking over snow, Hazerswoude, The Netherlands, February
Brown Bear (Ursus arctos) mother and 3-4 month cubs foraging for clams. Katmai National Park, Alaska, USA, June
