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Common swifts (Apus apus) flying overhead, Wiltshire, UK, June. Digital composite image
Starling murmuration, hundreds of thousands of starlings (Sturnus vulgaris) gather in the skies near their night time roost. Gretna Green, Scotland, January
Red kite (Milvus milvus) in flight in the snow, Wales, February. Bookplate from Danny Greens The Long Journey North
Bittern (Botaurus stellaris) in flight, Minsmere RSPB reserve, Suffolk, UK, August
Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Red Kite (Milvus milvus) in flight. Wales, UK. November
Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
Northern / Hen Harrier (Circus cyaneus) and Kestrel (Falco tinnunculus) below, fighting in flight. Vosges, France, February
Snowy owl (Bubo scandiaca) female in flight. Canada. February
Three Red Kites (Milvus milvus) chasing each other in flight. Wales, UK. November
Barn owl (Tyto alba) flying in snow covered countryside, Surrey, England, UK, January. Digital composite
Gannet (Morus bassanus) adult turns in the air as it begins to plunge dive. Shetland Islands, Scotland, UK, July
White-tailed eagle (Haliaeetus albicilla) with backlit wings, and water spray, Norway, October
Budgerigars (Melopsittacus undulatus) flocking to find water, Northern Territory, Australia
Whooper swans (Cygnus cygnus), flying at sunset, Caerlaverock Wildfowl & Wetland Trust WWT, Dumfries & Galloway, Scotland, November 2016
Swift (Apus apus) flock screaming in flight against blue sky and clouds, Monmouthshire, Wales, UK, July
Chilean flamingos (Phoenicopterus chilensis) in flight over mountain peaks with glacier in the distance, Torres Del Paine National Park, Chile
Wandering albatross {Diomedea exulans} flying over open ocean, South Atlantic
Atlantic puffin (Fratercula arctica) landing, wings outstretched. Machias Seal Island, off the coast of Maine, USA. July
Barn owl (Tyto alba) in flight, Norfolk, England, UK, February
Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK. January
Peregrine falcon (Falco peregrinus) adult landing at nest with chicks, Vaala, Finland, June
Peregrine falcon (Falco peregrinus) in flight over cotton grass flowers, Vaala, Finland, June
Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive /Snowy Owl (Nyctea scandiaca) adult female flying over snow, winter, Europe Captive/ Falconers bird, Austria
Griffon vulture (Gyps fulvus) in flight, Monfrague National Park, Extremadura, Spain, March 2009
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Red kite (Milvus milvus) in flight, Gigrin Farm, Mid Wales, UK, March. Non-ex
Curlews (Numenius arquata) group flying over the sea during storm. Wales, UK December. Highly commended, Habitat division, British Wildlife Photography Awards (BWPA) competition 2012
Barn owl (Tyto alba) in flight. Suffolk, UK. June
Buzzard (Buteo buteo) in flight, portrait, Marlborough Downs, Wiltshire, UK. December
Pipistrelle bat (Pipistrellus pipistrellus) flying low over water. Surrey, England, August
Large flock of Pink-footed geese (Anser brachyrynchus) flying from overnight roost at dawn, The Wash, Snettisham, Norfolk, UK, January
Siberian flying squirrel (Pteromys volans orii) flying directly over photographer. Hokkaido, Japan. February
European roller (Coracias garrulus) in flight, Pusztaszer, Hungary, May
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Stellers sea-eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan, February
Female Large white / Cabbage white butterfly (Pieris brassicae) in flight, Surrey, UK. Multiple exposure
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Great Grey Owl (Strix nebulosa) hunting over snow, Kuhmo Finland, March
Kestrel (Falco tinnunculus) male hovering overhead, Cornwall, UK, April
Barn Swallow (Hirundo rustica) in flight. Portugal, Europe
Whooper swans (Cygnus cygnus) three coming into land, Hokkaido, Japan, February
Osprey (Pandion haliaetus) in flight catching a fish, Finland, July
Corys shearwater (Calonectris diomedea) in flight over sea, Pico, Azores, Portugal, June 2009
Arctic tern (Sterna paradisaea) in flight, Inner Farne, Farne Islands, Northumberland, June
Mallard (Anas platyrhynchos) male flying low over water. Cambridgeshire Fens, England, March
Hen harrier (Circus cyaneus) female hovering over moorland, Glen Tanar Estate, Deeside, Scotland, UK, June
Stellers sea-eagle (Haliaeetus pelagicus) landing on pack ice, Hokkaido, Japan, February
Barn owl (Tyto alba) in flight, captive, Cumbria, UK
Kingfisher (Alcedo atthis) taking off from water with caught fish. Worcestershire, UK, March
Booted Racket-tail hummingbird (Ocreatus underwoodii) male flying, feeding from flower, Mindo, Ecuador, Andes, South America, January
Arctic tern (Sterna paradisaea) hovering above the ocean, Svalbard, Norway June
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
RF- Carrion crow (Corvus corone) flying from a snow covered pine tree in a winter landscape. Valsavarenche, Gran Paradiso NP, Italy, March
Three Whooper swans (Cygnus cygnus) in flight, Lake Tysslingen, Sweden, March 2009 WWE BOOK. WWE INDOOR EXHIBITION
Little Owl (Athene noctua) taking flight from a post. Wales, UK, June
Oystercatchers (Haematopus ostralegus) arriving to roost at Snettisham on the Wash. Norfolk, January
Golden eagle (Aquila chrysaetos) sub-adult flying, Strathdearn, Inverness-shire, Scotland, UK, August
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Eurasian crag martin (Ptyonoprogne rupestris) Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May
White chinned petrel (Procellaria aequinoctialis) in flight low over the water off Kaikoura, Canterbury, New Zealand
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Small Tortoiseshell Butterfly (Aglais urticae) feeding on scabious flowers in hay meadow, Wensum Valley, Norfolk, England, UK, July
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Grizzly bear (Ursus arctos horribilis) and Grey wolf (Canis lupus) on beach, Katmai National Park, Alaska, USA, August
White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
Lapwings (Vanellus vanellus), Golden plover (Pluvialis apricaria), Wigeon (Anas penelope) and Common Teal (Anas crecca) flying over flooded marshes in winter, with a rainbow in the background
Flock of Manx shearwaters (Puffinus puffinus) in flight over sea, near Ardnamurchan Point, Inner Hebrides, Scotland, UK, July 2011
Hen harrier (Circus cyaneus) adult female landing on rock in moorland habitat, Glen Tanar Estate, Grampian, Scotland, UK, June
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Northern gannet (Morus bassanus) colony with low clouds over cliff top, St Kilda, Scotland, May 2009
Common starling (Sturnus vulgaris) flocking prior to roosting, Scotland, January 2009
Golden eagle (Aquila chrysaetus) Kuusamo, Finland, January
Golden eagle (Aquila chrysaetos) adult female taking off, flying over mountain landscape, trained bird photographed during filming in Glenveagh National Park, Donegal, Ireland. August 2010
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
