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Carp (Cyprinus carpio) Aare River, Switzerland 2006
Tompot blenny (Parablennius gattorugine) in bright summer mating colours, peering out from his home in a discarded tube, where he was guarding his eggs (not visible in this photo), Swanage Pier
Killer Whales / Orcas (Orcinus orca)Killer whales / Orcas (Orcinus orca) large pod including calf traveling together while foraging on large schools of Herring (Clupea harengus) in the cold waters of northern Norway, January
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef. Whilst a Predatory bandcheek wrasse (Oxycheilinus digrammus)
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
Female Anglerfish fanfin seadevil (Caulophryne jordani) - Deep sea species from Atlantic Ocean, found at 700-3000m
Leafy seadragon (Phycodurus eques) male carrying eggs, swims over seagrass meadow. Wool Bay, Edithburgh, South Australia. Gulf of St Vincent
Northern pike (Esox lucius) amongst Shining pondweed (Potamogeton lucens). Lake Neuchatel, Switzerland, September. Photographed for The Freshwater Project
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
School of Blackfin barracuda (Sphyraena qenie) forming circle in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Weedy seadragon (Phyllopteryx taeniolatus) male carries eggs through a kelp forest (Macrocystis pyrifera) in Tasmania, Australia. Tasmania is the only part of Australia with giant kelp forests
Inflated spotted porcupinefish (Diodon hystrix), Maui, Hawaii
Portrait of Ballan wrasse (Labrus bergylta), St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, October 2011
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Bat fish juvenile {Halicutaea sp} deep sea
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Starry smooth-hound shark (Mustelus asterias) resting on seabed, Channel Isles, UK
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Brown trout (Salmo trutta) leaping for a damselfly. (digitally enhanced)
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
Spotted Wobbegong Shark (Orectolobus maculatus) lying in seaweed. Manly, Sydney, New South Wales, Australia. Pacific Ocean. November
Atlantic halibut (Hippoglossus hippoglossus) portrait, Saltstraumen, Bod, Norway, October 2008
Kingfisher (Alcedo atthis) perched with fish in beak. Worcestershire, UK, September
A portrait of a Mediterranean parrotfish (Sparisoma cretense) above an algae covered rock. El Hierro, Canary Islands, Spain. East Atlantic Ocean
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Great white shark (Carcharodon carcharias) underwater, Guadalupe Island, Mexico (North Pacific)
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Illuminated netdevil (Linophyne arboritera), male anglerfish attached to much larger female, captive from deep sea
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Lined seahorses (Hippocampus erectus) amongst corals, The Bahamas
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Brown trout (Salmo trutta) in shallow water migrating upstream, Bornholm, Denmark, November 2008
Bream (Abramis brama) and European chub (Leuciscus / Squalius cephalus) shoal in a calm part of the Rhine, Schaffhausen, Switzerland, February 2009 WWE OUTDOOR EXHIBITION
Puffin (Fraterclua arctica) with fish in beak, Saltee Island, Ireland June 2009
Pike (Esox lucius) in disused quarry, Stoney Stanton, Stoney Cove, Leicestershire, UK, June
Kingfisher (Alcedo atthis) taking off from water with caught fish. Worcestershire, UK, March