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Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October
Konik horses (Equus caballus) - wild Konik filly, Millingerwaard nature reserve, Netherlands, April
Andalusian mare running in snow storm, Colorado, USA
Wild horse / mustang, mare rolling, Pryor Mountains, Montana, USA, July
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Proboscis Monkey (Nasalis larvatus) female sitting in tree, partially obscured by soft focus leaves in foreground, Bako National Park, Sarawak, Borneo, Malaysia
Polar bears (Ursus maritimus) female with cubs aged two years travelling along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
An adult female California sea lion (Zalophus californianus) head portrait, in a rocky underwater cave. La Paz, Baja California Mexico. Sea of Cortez, East Pacific Ocean. October
Crested Newt (Triturus cristatus carnifex) underwater view of female swimming on surface, Traversari lake, Campigna National Park, Emilia Romagna and Tuscany regions, Italy
Polar bear (Ursus maritimus) sow plays with her spring cub outside their den in late winter, Alaska
Jaguar (Panthera onca) female, hiding in dense high grass. Cuiaba river, Pantanal, Mato Grosso do Sul Province. Brazil, South America
A wild Misaki-uma breeding stallion (right) brings back one of his mares to his band while another breeding stallion (left) charges him away, in the Cape Toi Reserve, Miyazaki Prefecture
Humpback whale (Megaptera novaeangliae) mother and calf swimming, Hawaii, Pacific Ocean
Japanese macaque (Macaca fuscata) holding young while bathing in hot spring to keep warm, only females and young bathe, winter, Jigokudani, Joshinetsu Kogen NP, Nagano, Japan
Female Japanese macaques (Macaca fuscata) in a hot spring to keep warm, only females and young bathe, Jigokudani, Joshinetsu Kogen NP, Nagano, Japan
Japanese macaques (Macaca fuscata) warming up by bathing in a hot spring during snowfall, only females and young bathe, Jigokudani, Joshinetsu Kogen NP, Nagano, Japan
Common blue butterfly (Polyommatus icarus) females awaking at sunrise among Harebells (Campanula rotundifolia) Surrey, England, digital composite
Verreauxs sifaka (Propithecus verreauxi) running with young on back, Berenty Reserve, Madagascar
Japanese macaque / Snow monkey {Macaca fuscata} adult female bathing in hot springs tapping the water surface with hand in play, water at 40 degrees, Jigokudani, Nagano, Japan
Red lechwe pair running & jumping in swamp {Kobus leche}. Khwai river, Moremi GR, Botswana
Female European Brown bear with cubs following {Ursus arctos} Lapland Finland
Female Roe deer {Capreolus capreolus} head portrait in flowering field, England, UK, Europe
Southern elephant seal {Mirounga leonina} female defensive threat display, Valdez, Argentina