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Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Female Brown bear (Ursus arctos) with two cubs, eating fish in lake Kuril, Kronotsky Nature Reserve, Kamchatka, Russia
Female Brown bear (Ursus arctos) with cub playing in lake Kuril, Kronotsky Nature Reserve, Kamchatka, Russia
Siberian tiger (Panthera tigris altaica) at night, taken with remote camera in Land of the Leopard National Park, Far East Russia
Underwater view of Brown bear (Ursus arctos) paw fishing for Sockeye salmon (Oncorhynchus nerka) with paw outstretched. Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August
Brown bear (Ursus arctos) walking across rock in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. September
Sika deer (Cervus nippon) doe standing on rocky outcrop overlooking mountain forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. January
Sika deer (Cervus nippon) stag wading through river in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) herd feeding beside pool in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) doe with fawn standing on rocky outcrop overlooking forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Asian black bear (Ursus thibetanus) scent marking tree by rubbing back on it, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it, Land of the Leopard National Park, Russian Far East. Endangered
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Amur leopard (Panthera pardus orientalis) walking past snow-covered trees on mountain slope, Land of the Leopard National Park, Russian Far East. Critically endangered. Taken with remote camera
Sika deer (Cervus nippon) doe walking across snow covered mountain slope, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Brown bear (Ursus arctos), portrait, Kronotsky Nature Reserve, Kamchatka, Russia
Steller sea lions (Eumetopias jubatus) swimming in the Bering Sea near Verkhoturova Island, Russia
Peschanaya Bay, Medny Island, Commander Islands, Russia
Brown bear (Ursus arctos) seen from underwater, fishing for Sockeye salmon, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
Brown bear (Ursus arctos) portrait, whilst fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
Brown bear (Ursus arctos), fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August