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Five Cavalier King Charles Spaniel sitting, black-and-tan, tricolour, blenheim and ruby coated, sitting in autumn leaves
Snow leopard (Panthera uncia) portrait with ears back. Captive
Portrait of English Springer Spaniel (field type). Elkhorn, Wisconsin, USA, January
Orangutan baby (Pongo pygmaeus) head portrait of baby, Semengoh Nature reserve, Sarawak, Borneo, Malaysia. Endangered
Portrait of a young Little Owl (Athene noctua)
Brown capuchin monkey (Sapajus macrocephalus). displaying submissive behavior. Captive, Peru
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Portrait of rabbit in leaf litter, Okunoshima Rabbit Island, Takehara, Hiroshima, Japan
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Tricolour Border Collie puppy
Blackbird (Turdus merula) male eating winfall apples in snow, Broxwater, Cornwall, UK. December
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
Arctic fox (Vulpes lagopus) portrait of juvenile, winter pelage. Dovrefjell National Park, Norway. September
A Florida manatee (Trichechus manatus latirostrus) head portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
Norfolk terrier portrait, lying in autumn foliage
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Three Sockeye salmon/ Red salmon (Oncorhynchus nerka) swimming upstream as they migrate back to the river of their birth to spawn
Border collie x Bearded collie puppy, aged 4 months, sitting looking up with tongue out, portrait
Basking shark (Cetorhinus maximus) feeding on plankton in surface waters close to the island of Coll, Inner Hebrides. Scotland, UK. Atlantic Ocean. June
Close up of a silverback Mountain gorilla (Gorilla beringei beringei) face with eyes closed, and hand / fingers, Virunga National Park, Democratic Republic of Congo
Frill-necked lizard (Chlamydosaurus kingii) threat display with neck frill extended, Cooktown, Queensland, Australia
Male Mantled howler monkey (Alouatta palliata) on foot bridge, La Selva Biological Station, Costa Rica
Grey seal (Halichoerus grypus) pup waving, UK
Snow leopard (Panthera uncia) with ears back. Captive
Eurasian lynx (Lynx lynx) yawning. Captive
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands
Deckens sifaka (Propithecus deckenii) grooming each other, Tsimembo area, Madagascar. Lenses for Conservation project
Three Little owls (Athene noctua) looking out of a nest hole, Cumbria, UK, August. Captive
Yorkshire Terrier mother, Evie, with her two Yorkishire terrier x Poodle Yorkipoo pups, age 7 weeks
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Caribbean reef squid (Sepioteuthis sepioidea) swimming in open water at night, Guadeloupe Island, Mexico. Caribbean
Mareeba rock wallaby (Petrogale mareeba), female feeding on leaf. Granite Gorge Nature Park, near Mareeba, North Queensland, Australia
Arctic fox (Vulpes lagopus), juvenile licking lips, winter pelage. Dovrefjell National Park, Norway. September
Burrowing owl (Athene cunicularia) rotating head, Arizona, USA
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Indonesian blue-tongued skink (Tiliqua gigas) with tongue extended, captive, from Papua New Guinea
Wolf-eel (Anarrhichthys ocellatus) Vancouver Island, British Columbia, Pacific Ocean
Sperm whale (Physeter macrocephalus) at surface, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Lion (Panthera leo) close up of teeth while its snarling, captive
Bichon frise beside Colorado blue columbine (Aquilegia coerulea) flowers. Old Lyme, Connecticut, USA. May
Bichon frise standing on rock amongst Rhododendron flowers. Haddam, Connecticut, USA. May
RF- Yorkipoo dog, Yorkshire terrier cross Poodle, Oscar, age 6 months
Border Collie puppy, 13 weeks
Cavalier King Charles Spaniel, two puppies on grass, blenheim and tricolour, 5 weeks
Caracal (Caracal caracal) male, standing on rock against blue sky, Spain. Captive, occurs in Africa and Asia
Cockapoo puppy, aged 18 weeks, running, portrait
Sperm whale (Physeter macrocephalus) with fully open mouth, Dominica, Caribbean Sea, Atlantic Ocean. Photo taken under permit
Black Welsh mountain sheep lamb calling. Wales UK, March
Two male Polar bears (Ursus maritimus) standing on hind legs, sparring, Churchill, Canada. November
Female Red fox (Vulpes vulpes) walking along tree trunk in heavy fog at night. Light source behind the subject creates dramatic, volumetric lighting, Hungary, January
Black moray (Muraena augusti) and Tiger moray eels (Enchelycore anatina) peering out from a rock crevice with mouths open, Tenerife, Canary Islands, Atlantic Ocean
RF- Portrait of male Parsons Chameleon (Chamaeleo parsonii), Ranomafana National Park, South eastern Madagascar
Spotted porcupinefish (Diodon hystrix), inflated with seawater, a defensive behaviour, Hawaii, Pacific Ocean
Whale shark (Rhincodon typus) feeding close to fishermens boats in the early morning, Triton Bay, West Papua, Indonesia, Pacific Ocean
Galapagos land iguana (Conolophus subcristatus) recently reintroduced to Santiago island where it had become extinct, feeding on Yellow cordia (Cordia lutea) leaves, Santiago Island, Galapagos
Fox (Vulpes vulpes) drinking water from a sauna pool in a garden; low angle view looking up through water, Verteskozma, Hungary
Red fox in a snow storm (Vulpes vulpes) Winter winds on Kamchatka can reach 40 meters a second. Kronotskty Zapovednik, Kamchatka, Russia
Yellow bellied snapping turtle (Elseya flaviventralis) about to surface for air, Arnhem Escarpment Creek, Northern Territory, Australia. June
Sandstone long-necked turtle (Chelodina burrungandjii) actively foraging at night, Pentecost River, Western Australia. April
Common tern (Sterna hirundo) flying low over water. Hjalstaviken, Uppland, Sweden
Yacare Caiman (Caiman yacare) gaping to regulate its body temperature, with attendant hoverflies, the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Ezo red fox (Vulpes vulpes schrencki) sitting on snow, licking lips after eating Vole. Hokkaido, Japan. April
Yellow pygmy goby (Lubricogobius exiguus) portrait, Kashiwajima, Japan
Ornamental koi carp (Cyprinus carpio) group with open mouths at lake surface. Xishuangbanna Tropical and Flowers Garden, Jinghong, Yunnan, China
Umbilicate woodcock orchid (Ophrys umbilicata) flowers. Cyprus. April
Mining bee / Little flower bee (Anthophora bimaculata) taking nectar and pollen from Common fleabane (Pulicaria dysenterica) flower, with pollen covered scopa, Surrey, England, UK. August
Chimpanzee juvenile (Pan troglodytes schweinfurthii) in a tree. Kibale National Park, Uganda. January
Tricolour merle Collie puppy, Indie, age 10 weeks
Wood warbler (Phylloscopus sibilatrix) singing whilst perched on branch, silhouetted at dawn. Sheffield, England, UK. May
Culpea or Andean fox (Pseudalopex culpaeus) at rest, vicinity of Laguna Verde, Reserva Eduardo Avaroa, altiplano, Bolivia
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
Head portrait of grey Andalusian stallion in dark barn, Northern France, Europe
Masked tree frog (Smilisca phaeota) El Arenal region, Costa Rica
Crowned sifaka (Propithecus coronatus), Katsepy, Madagascar
Nile crocodile (Crocodylus niloticus). Zimanga Private Game Reserve, KwaZulu-Natal, South Africa. September
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
RF - Sumatran tiger (Panthera tigris altaica / Panthera tigris tigris) staring at camera, licking nose, captive
Shetland sheepdog running in Common bluet (Houstonia caerulea) field. Haddam, Connecticut, USA. May
Cavalier King Charles Spaniel x Poodle Cavapoo and red toy Poodle
Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp)Grizzly bear (Ursus arctos) standing at top of waterfall catching Salmon (Salmo sp.), Brooks Falls, Katmai National Park, Alaska. July
Grizzly bear (Ursus arctos) female with cub seeking attention from its mother, Lake Clark National Park, Silver Salmon, Alaska. July
Polar bear (Ursus maritimus) female, resting on ice with cub sitting next to her waving and poking out tongue, Svalbard, Norway. April
Yacare caiman (Caiman yacare) with mouth open, head portrait, Cuiaba River, Pantanal, Mato Grosso, Brazil
Bluethroat (Luscinia svecica cyanecula) male, perched on tree stump singing, Germany. March